The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
In Germany, the two forms have long been popularly distinguished, the
former by the name of “Steinkrebs,” {290} or “stone crayfish,” and
the latter by that of “Edelkrebs,” or “noble crayfish.”
Milne Edwards, it will be observed, speaks of these two forms of
crayfish as “varieties” of the species _Astacus fluviatilis_; but,
even as far back as the year 1803 some zoologists began to regard the
“stone crayfish” as a distinct species, to which Schrank applied the
name of _Astacus torrentium_, while the “noble crayfish” remained
in possession of the old denomination, _Astacus fluviatilis_;
and, subsequently, various forms of “stone-crayfishes” have been
further distinguished as the species _Astacus saxatilis_, _A.
tristis_, _A. pallipes_, _A. fontinalis_, &c. On the other hand,
Dr. Gerstfeldt,[19] who has devoted especial attention to the
question, denies that these are anything more than varieties of one
species; but he holds this and Milne Edwards’s “second variety” to be
specifically distinct from one another.
We thus find ourselves in the presence of three views respecting the
English and French crayfishes.
1. They are all varieties of one species—_A. fluviatilis_.
2. There are two species—_A. fluviatilis_, and _A. torrentium_, of
which last there are several varieties.
3. There are, at fewest, five or six distinct species.
Before adopting the one or the other of these views, it is necessary
to form a definite conception of the meaning of the terms “species”
and “variety.” {291}
[19] “Ueber die Flusskrebse Europas.” Mém. de l’Acad. de St.
Petersburg, 1859.
The word “species” in Biology has two significations; the one based
upon morphological, the other upon physiological considerations.
A species, in the strictly morphological sense, is simply an
assemblage of individuals which agree with one another, and differ
from the rest of the living world in the sum of their morphological
characters; that is to say, in the structure and in the development
of both sexes. If the sum of these characters in one group is
represented by A, and that in another by A + _n_; the two are
morphological species, whether _n_ represents an important or an
unimportant difference.
The great majority of species described in works on Systematic
Zoology are merely morphological species. That is to say, one or more
specimens of a kind of animal having been obtained, these specimens
have been found to differ from any previously known by the character
or characters _n_; and this difference constitutes the definition of
the new species, and is all we really know about its distinctness.
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