The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
The case is altered when we reach the Middle Lias. In fact this
yields several forms of a genus, _Eryma_ (fig. 80, B), which also
occurs in the overlying strata almost up to the top of the Jurassic
series, and presents so many variations that nearly forty different
species have been recognised. _Eryma_ is, in all respects, an
Astacomorph, and so far as can be seen, it differs from the existing
genera only in such respects as those in which they differ from one
another. Thus it is quite certain that Astacomorphous _Crustacea_
have existed since a period so remote as the older part of the
Mesozoic period; and any hesitation in admitting this singular
persistency of type on the part of the crayfishes, is at once removed
by the consideration of the fact that, along with _Eryma_, in the
Middle Lias, prawn-like _Crustacea_, generically identical with
the existing _Penæus_, flourished in the sea {342} and left their
remains in the mud of the ancient sea bottom.
[Illustration: FIG. 81.—_Hoploparia longimana_ (2/3 nat. size).—_cp_,
carapace; _r_, rostrum, T, telson; XV., XVI., first and second
abdominal somites; 10, forceps; 20, last abdominal appendage.]
_Eryma_ is the only crustacean, which can be certainly ascribed to
the _Astacomorpha_, that has hitherto been found in the strata from
the Middle Lias to the lithographic slates; which last lie in the
upper part of the Jurassic series. In the freshwater beds of the
Wealden, no _Astacomorpha_ are known, and although no very great
weight is to be attached to a negative fact of this kind, it is, so
far, evidence that the _Astacomorpha_ had not yet taken to freshwater
life. In the marine deposits of the Cretaceous epoch, however,
astacomorphous forms, which {343} are known by the generic names of
_Hoploparia_ and _Enoploclytia_, are abundant.
The differences between these two genera, and between both and
_Eryma_, are altogether insignificant from a broad morphological
point of view. They appear to me to be of less importance than those
which obtain between the different existing genera of crayfishes.
_Hoploparia_ is found in the London clay. It therefore extends beyond
the bounds of the Mesozoic epoch into the older Tertiary. But when
this genus is compared with the existing _Homarus_ and _Nephrops_,
it is found partly to resemble the one and partly the other. Thus,
on one line, the actual series of forms which have succeeded one
another from the Liassic epoch to the present day, is such as must
have existed if the common lobster and the Norway lobster are the
descendants of _Erymoid_ crustaceans which inhabited the seas of the
Liassic epoch.
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