The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
The free extremity of the eyestalk presents a convex, soft, and
transparent surface, limited by an oval contour. The cuticle in
this region, which is termed the _cornea_, (fig. 28, _a_), is, in
fact, somewhat thinner and less distinctly laminated than in the
rest of the eyestalk, and it contains no calcareous matter. But
it is directly continuous with the rest of the exoskeleton of the
eyestalk, to which it stands in somewhat the same relation as the
soft integument of an articulation does to the adjacent hard parts.
[Illustration: FIG. 28.—_Astacus fluviatilis._—A, a vertical section
of the eye-stalk (× 6); B, a small portion of the same, showing the
visual apparatus more highly magnified; _a_, cornea; _b_, outer dark
zone; _c_, outer white zone; _d_, middle dark zone; _e_, inner white
zone; _f_, inner dark zone; _cr_, crystalline cones; _g_, optic
ganglion; _op_, optic nerve; _sp_, striated spindles.]
The _cornea_ is divided into a great number of minute, usually square
facets, by faint lines, which cross it from side {119} to side
nearly at right angles with one another. A longitudinal section shows
that both the horizontal and the vertical contours of the cornea
are very nearly semicircular, and that the lines which mark off the
facets merely arise from a slight modification of its substance
between the facets. The outer contour of each facet forms part of
the general curvature of the outer face of the cornea; the inner
contour sometimes exhibits a slight deviation {120} from the general
curvature of the inner face, but usually nearly coincides with it.
When a longitudinal or a transverse section is taken through the
whole eyestalk, the optic nerve (fig. 28, A, _op_) is seen to
traverse its centre. At first narrow and cylindrical, it expands
towards its extremity into a sort of bulb (B, _g_), the outer surface
of which is curved in correspondence with the inner surface of the
cornea. The terminal half of the bulb contains a great quantity of
dark colouring matter or pigment, and, in section, appears as what
may be termed the _inner dark zone_ (_f_). Outside this, and in
connection with it, follows a white line, the _inner white zone_
(_e_), then comes a _middle dark zone_ (_d_); outside this an outer
pale band, which may be called the _outer white zone_ (_c_), and
between this and the cornea (_a_) is another broad band of dark
pigment, the _outer dark zone_ (_b_).
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