The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
But, in the first place, there is no evidence, nor any probability,
that there is anything corresponding to a retina on the outer face of
the crystalline cone; and secondly, if there were, it is incredible
that, with such an arrangement of the refractive media as exists in
the cornea and crystalline cones, rays proceeding from points in
the external world should be brought to a focus in correspondingly
related points of the surface of the supposed retina. But without
this no picture could be formed, and no distinct vision could take
place. It is very probable, therefore, that the visual pyramids do
not play the part of the simple eyes of the _Vertebrata_, and the
only alternative appears to be the adoption of a modification of the
theory of _mosaic vision_, propounded many years by Johannes Müller.
{123}
Each visual pyramid, isolated from its fellows by its coat of
pigment, may be supposed, in fact, to play the part of a very narrow
straight tube, with blackened walls, one end of which is turned
towards the external world, while the other incloses the extremity of
one of the nerve fibres. The only light which can reach the latter,
under these circumstances, is such as proceeds from points which lie
in the direction of a straight line represented by the produced axis
of the tubes.
[Illustration: FIG. 29.—Diagram showing the course of rays of light
from three points _x_, _y_, _z_, through the nine visual rods
(supposed to be empty tubes) A–I of a compound eye; _a–i_, the nerve
fibres connected with the visual rods.]
Suppose A–I to be nine such tubes, _a–i_ the corresponding nerve
fibres, and _x_ _y_ _z_ three points from which light proceeds. Then
it will be obvious that the only light {124} from _x_ which will
excite sensation, will be the ray which traverses B and reaches the
nerve-fibre _b_, while that from _y_ will affect only _e_, and that
from _x_ only _h_. The result, translated into sensation, will be
three points of light on a dark ground, each of which answers to one
of the luminous points, and indicates its direction in reference to
the eye and its angular distance from the other two.[10]
The only modification needed in the original form of the theory of
mosaic vision, is the supposition that part, or the whole, of the
visual rod, is not merely a passive transmitter of light to the
nerve-fibre, but is, itself, in someway concerned in transmuting
the mode of motion, light, into that other mode of motion which we
call nervous energy. The visual rod is, in fact, to be regarded as
the physiological end of the nerve, and the instrument by which the
conversion of the one form of motion into the other takes place; just
as the auditory hairs are instruments by which the sonorous waves are
converted into molecular movements of the substance of the auditory
nerves.
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