The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
To apply all this to the crayfish. It is, in a sense a house with
a great variety of rooms and offices, in which the work of the
indwelling life in feeding, breathing, moving, and reproducing
itself, is done. But the {140} same may be said of the crayfish’s
neighbours, the perch and the water-snail; and they do all these
things neither better nor worse, in relation to the conditions
of their existence, than the crayfish does. Yet the most cursory
inspection is sufficient to show that the “styles of architecture”
of the three are even more widely different than are those of the
Gothic, Italian, and Queen Anne houses.
That which Architecture, as an art conversant with pure form, is to
buildings, Morphology, as a science conversant with pure form, is
to animals and plants. And we may now proceed to occupy ourselves
exclusively with the morphological aspect of the crayfish.
* * * * *
As I have already mentioned, when dealing with the physiology of the
crayfish, the entire body of the animal, when reduced to its simplest
morphological expression, may be represented as a cylinder, closed
at each end, except so far as it is perforated by the alimentary
apertures (fig. 6); or we may say that it is a tube, inclosing
another tube, the edges of the two being continuous at their
extremities. The outer tube has a chitinous outer coat or cuticle,
which is continued on to the inner face of the inner tube. Neglecting
this for the present, the outermost part of the wall of the outer
tube, which answers to the _epidermis_ of the higher animals, and
the innermost part of the wall of the inner tube, which is an
_epithelium_, are formed by a layer of nucleated cells. A continuous
layer of cells, therefore, is everywhere to {141} be found on both
the external and the internal free surfaces of the body. So far as
these cells belong to the proper external wall of the body, they
constitute the _ectoderm_, and so far as they belong to its proper
internal wall, they compose the _endoderm_. Between these two layers
of nucleated cells lie all the other parts of the body, composed of
connective tissue, muscles, vessels, and nerves; and all these (with
the exception of the ganglionic chain, which we shall see properly
belongs to the ectoderm) may be regarded as a single thick stratum,
which, as it lies between the ectoderm and the endoderm, is called
the _mesoderm_.
If the intestine were closed posteriorly instead of opening by the
vent, the crayfish would virtually be an elongated sac, with one
opening, the mouth, affording an entrance into the alimentary cavity:
and, round this cavity, the three layers just referred to—endoderm,
mesoderm, and ectoderm—would be disposed concentrically.
* * * * *
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