Just above the smaller eye there lies a mass of cells of peculiar
structure (Fig. 69, _nc_). They are of a rounded polygonal contour, with
a comparatively small circular nucleus in the centre, and are found in
this region only. In and amongst them bundles of fibrous tissue are found
in the sections, which pass from the surface cells to the supporting
lamella. Claus describes the contents of these cells as coarsely granular
protoplasm and says they cannot be taken for ganglion cells. He is
inclined to believe that they play the part of a special supporting
tissue. Schewiakoff, on the other hand, is convinced that they are
ganglion cells, and finds processes passing out from them (’89, Taf. II,
Fig. 22). I find, however, that the cell contours are perfectly regular
and clearly without processes, and it is incomprehensible to me how,
if his material was at all well preserved, he could for a moment have
taken them for the same thing as the big multipolar ganglion cells with
large nucleus and nucleolus which lie in about the same region and were
correctly described and figured by Claus but are not specially mentioned
by Schewiakoff. I cannot agree with Claus, however, that their contents
are composed of coarsely granular protoplasm. That which appears such
by low magnification shows itself under high powers to be a beautiful
network with thickenings at the nodes of the meshes, which is brought
out very plainly by a cytoplasmic stain such as Lyons blue. Around the
nucleus is seen a more or less well-defined clear zone. What the function
of the cell is remains as unknown to me as to Claus and Schewiakoff.
There is left one more point in reference to the nervous system upon
which I wish to say a word. Claus and Schewiakoff both describe the wall
of the crystalline sac as structureless, formed by the bare supporting
lamella. The credit is due to H. V. Wilson of finding in Chiropsalmus
that it has a special lining of epithelial cells, which he figures as
a continuous, flattened layer. In both Charybdea and Tripedalia I find
traces of the same in nuclei here and there, but whether they are the
remains of a once continuous layer or not the sections do not show
satisfactorily.
This ends the account of what it seemed worth while to say at present
upon the nervous system. In concluding, the writer wishes to express
his thanks for the help afforded by Dr. Wilson’s notes, in particular
on the subject of the vascular lamellæ, and desires to make especial
acknowledgment of his indebtedness to Professor Brooks, whose
suggestions, based upon many years of experience with the Medusæ,
have been most welcome and helpful, and whose evidences of unfailing
kindliness, both in Jamaica at the time the material was obtained and
in Baltimore when it was being studied in the laboratory, take a most
honored part in the pleasant memories associated with the work.
LITERATURE REFERRED TO.
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