The Descent of Man and Selection in Relation to Sex, Vol. IDarwin, Charles
Science
The Descent of Man and Selection in Relation to Sex, Vol. I
Darwin, Charles
Evolution (Biology); Human beings -- Origin; Sex differences; Sexual dimorphism (Animals); Sexual selection in animals
_Stridulating organs._—Beetles belonging to many and widely distinct
families possess these organs. The sound can sometimes be heard at the
distance of several feet or even yards,[494] but is not comparable with
that produced by the Orthoptera. The part which may be called the rasp
generally consists of a narrow slightly-raised surface, crossed by very
fine, parallel ribs, sometimes so fine as to cause iridescent colours,
and having a very elegant appearance under the microscope. In some
cases, for instance, with Typhæus, it could be plainly seen that
extremely minute, bristly, scale-like prominences, which cover the whole
surrounding surface in approximately parallel lines, give rise to the
ribs of the rasp by becoming confluent and straight, and at the same
time more prominent and smooth. A hard ridge on any adjoining part of
the body, which in some cases is specially modified for the purpose,
serves as the scraper for the rasp. The scraper is rapidly moved across
the rasp, or conversely the rasp across the scraper.
[Illustration: Fig. 24. Necrophorus (from Landois). _r._ The two rasps.
Left-hand figure, part of the rasp highly magnified.]
These organs are situated in widely different positions. In the
carrion-beetles (Necrophorus) two parallel rasps (_r_, fig. 24) stand on
the dorsal surface of the fifth abdominal segment, each rasp being
crossed, as described by Landois,[495] by from 126 to 140 fine ribs.
These ribs are scraped by the posterior margins of the elytra, a small
portion of which projects beyond the general outline. In many
Crioceridæ, and in _Clythra 4-punctata_ (one of the Chrysomelidæ), and
in some Tenebrionidæ, &c.,[496] the rasp is seated on the dorsal apex of
the abdomen, on the pygidium or pro-pygidium, and is scraped as above by
the elytra. In Heterocerus, which belongs to another family, the rasps
are placed on the sides of the first abdominal segment, and are scraped
by ridges on the femora.[497] In certain Curculionidæ and Carabidæ,[498]
the parts are completely reversed in position, for the rasps are seated
on the inferior surface of the elytra, near their apices, or along their
outer margins, and the edges of the abdominal segments serve as the
scrapers. In _Pelobius hermanni_ (one of Dytiscidæ or water-beetles) a
strong ridge runs parallel and near to the sutural margin of the elytra,
and is crossed by ribs, coarse in the middle part, but becoming
gradually finer at both ends, especially at the upper end; when this
insect is held under water or in the air, a stridulating noise is
produced by scraping the extreme horny margin of the abdomen against the
rasp. In a great number of long-horned beetles (Longicornia) the organs
are altogether differently situated, the rasp being on the meso-thorax,
which is rubbed against the pro-thorax; Landois counted 238 very fine
ribs on the rasp of _Cerambyx heros_.
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