The Descent of Man and Selection in Relation to Sex, Vol. II (1st Edition)Darwin, Charles
Science
The Descent of Man and Selection in Relation to Sex, Vol. II (1st Edition)
Darwin, Charles
Evolution (Biology); Human beings -- Origin; Sex differences; Sexual dimorphism (Animals); Sexual selection in animals
The successive steps in the shading of the ball-and-socket ocelli can be
followed out with equal clearness. The brown, orange, and pale-leaden
narrow zones which border the lower black mark of the elliptic ornament
can be seen gradually to become more and more softened and shaded into
each other, with the upper lighter part towards the left-hand corner
rendered still lighter, so as to become almost white. But even in the
most perfect ball-and-socket ocelli a slight difference in the tints,
though not in the shading, between the upper and lower parts of the ball
can be perceived (as was before especially noticed), the line of
separation being oblique, in the same direction with the bright coloured
shades of the elliptic ornaments. Thus almost every minute detail in the
shape and colouring of the ball-and-socket ocelli can be shewn to follow
from gradual changes in the elliptic ornaments; and the development of
the latter can be traced by equally small steps from the union of two
almost simple spots, the lower one (fig. 57) having some dull fulvous
shading on the upper side.
[Illustration: Fig. 59. Portion near summit of one of the Secondary
wing-feathers, bearing perfect ball-and-socket ocelli.
_a._ Ornamented upper part.
_b._ Uppermost, imperfect ball-and-socket ocellus. (The
shading above the white mark on the summit of the ocellus is
here a little too dark.)
_c._ Perfect ocellus.]
The extremities of the longer secondary feathers which bear the perfect
ball-and-socket ocelli are peculiarly ornamented. (Fig. 59.) The oblique
longitudinal stripes suddenly cease upwards and become confused, and
above this limit the whole upper end of the feather (_a_) is covered
with white dots, surrounded by little black rings, standing on a dark
ground. Even the oblique stripe belonging to the uppermost ocellus (_b_)
is represented only by a very short irregular black mark with the usual,
curved, transverse base. As this stripe is thus abruptly cut off above,
we can understand, from what has gone before, how it is that the upper
thickened part of the ring is absent in the uppermost ocellus; for, as
before stated, this thickened part is apparently formed by a broken
prolongation of the next higher spot in the same row. From the absence
of the upper and thickened part of the ring, the uppermost ocellus,
though perfect in all other respects, appears as if its top had been
obliquely sliced off. It would, I think, perplex any one, who believes
that the plumage of the Argus pheasant was created as we now see it, to
account for the imperfect condition of the uppermost ocelli. I should
add that in the secondary wing-feather farthest from the body all the
ocelli are smaller and less perfect than on the other feathers, with
the upper parts of the external black rings deficient, as in the case
just mentioned. The imperfection here seems to be connected with the
fact that the spots on this feather shew less tendency than usual to
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