The Deseado Formation of PatagoniaLoomis, Frederic Brewster
Science
The Deseado Formation of Patagonia
Loomis, Frederic Brewster
Paleontology -- Patagonia (Argentina and Chile)
The treatment of these forms has been as varied as their sizes.
Ameghino, with his idea that the Casamayor and Deseado beds
were Cretaceous in age, groups the larger forms as a suborder,
_Sparassodonta_, and considers them ancestral to the _Creodonta_; while
the small forms make up his _Sarcobora_ which he considered ancestral
on one side to the rodents, on the other to the diprotodont marsupials.
Sinclair, after showing the marked similarity of the _Sparassodonta_
to the polyprotodont marsupials, especially the genus _Thylacynus_,
abandons that term and puts them in the family _Thylacynidae_ along
with the Australian forms; the _Microbiotheridae_ he finds similar to
opossums and puts in the family _Didelphidae_; while the remaining
small diprotodont forms he associates with _Caenolestes_, and using
Ameghino’s families as subfamilies makes three divisions of the family,
_Palaeothentinae_, _Garzoninae_, and _Abderitinae_. Matthew finds
the sparassodonts to be true marsupials, and without phylogenetic
relationship with the creodonts. Gregory diagrams the sparassodonts
as coming from generalized didelphids and derives them from the same
line as the Australian polyprotodonts; while the small caenolestoids
represent a line of descent from some still earlier generalized
polyprotodonts and a separate stem from the Australian diprotodonts.
Sinclair has had the most complete material on which to work, and
with his general grouping I have come to agree. This recognizes
three divisions of South American Marsupials, the _Didelphidae_,
representatives of which have not yet been found in the Deseado, though
occurring in both the earlier and later formations; the _Caenolestidae_
represented today by _Caenolestes_, the only survivor of the South
American diprotodonts; and the _Borhyaenidae_ (= _Thylacynidae_
of Sinclair this name having been used to indicate a much nearer
relationship to the Australian _Thylacynus_ than I feel is warranted),
which includes a large range of medium to large sized animals ranging
from the Casamayor formation throughout the Santa Cruz beds.
The locality from which these marsupials emigrated to South America
and the time of their arrival is not yet agreed upon, and can not
be settled until much more complete material is discovered in the
Casamayor formation. I feel, however, that the three groups were
separate when they entered South America.
=Borhyaenidae=
Ameghino has grouped in this family a considerable number of genera of
powerful, wolf-like carnivorous marsupials, characterized by a dental
formula
4-3 1 4 3
----------,[24]
3 1 4 3
Public-domain text, read in full here on John Shaqi.
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