The Deseado Formation of PatagoniaLoomis, Frederic Brewster
Science
The Deseado Formation of Patagonia
Loomis, Frederic Brewster
Paleontology -- Patagonia (Argentina and Chile)
as in _Adinotherium_, the chief difference being that the teeth are
less hypsodont than in the Santa Cruz genus. Little is known as yet
of the skeleton, but when more is known probably more distinctive
characters will appear. Ameghino made two species, _P. leptognathus_
which we also found, and _P. angustidens_ a much smaller form.
=Proadinotherium leptognathus= Ameghino
P. leptognathus Amegh., 1895, Bol. Inst. Geog. Argen., t. 15, p. 625.
P. leptognathus Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 467.
[Illustration: Fig. 81. Left lower jaw with incisor 3 and molar 2—½
natural size.]
Of this species we found on the Chico del Chubut River, west of Puerto
Visser, three specimens; the back of a skull as far forward as molar 2,
and two lower jaws. In general, the species is very similar, even to
size, to _Adinotherium ovinum_ of the Santa Cruz.
The upper molars are strongly hypsodont, curved teeth. On the upper
surface, the basin is subdivided by two strong cristae into three
smaller bays. In an early stage of wear, the second crista unites with
the posterior lobe, converting bay 3 into a pit. On the posterior
margin of the tooth, the cingulum is developed so as to appear like a
third crista, which inclosed bay 4, and when the tooth is worn, bay 4
becomes a pit also.
In my lower jaw incisor 3 is developed into a strong caniniform
tush. Most of the teeth are lacking, but lower molar 2 is a strongly
compressed, hypsodont tooth, surrounded by a thick layer of enamel.
This tooth rises 22 mm. above the well-developed roots, and is already
considerably worn down. The pillar is prominent as a strong fold in the
middle of the posterior crescent. In this specimen there is no trace
of the usual pit (3) indicative of the septum, but I should expect to
find it in a younger specimen. The mandible broadens in front into a
scoop-like anterior end, and the alveoli of the first two incisors
would indicate that they were proclivous. The alveoli for the other
teeth are arranged as in _Adinotherium_.
MEASUREMENTS
Skull, width across the zygomatic arches 148 mm.
Skull, width across opposite m. 3 (outside) 73 mm.
Upper dentition, molar 2, length 25 mm., width 13 mm.
Upper dentition, molar 3, length 23 mm., width 12 mm.
Lower dentition, incisor 3, length 13 mm., width 7 mm.
Lower dentition, molar 2, length 20 mm., width 7½ mm.
=Proadinotherium angustidens= Ameghino
P. angustidens Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 467.
This is based on a single lower tooth, which is considered either pm. 4
or m. 1, and measures 13 mm. long by 4½ mm. wide.
=Pronesodon= Ameghino
Pronesodon Amegh., 1895, Bol. Inst. Geog. Argen., t. 15, p. 626.
The genus is said to resemble _Proadinotherium_, but with the
caniniform incisors proportionally much shorter. An associated
calcaneum is shorter than that of _Adinotherium_ and longer than that
of _Nesodon_.
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