The Deseado Formation of PatagoniaLoomis, Frederic Brewster
Science
The Deseado Formation of Patagonia
Loomis, Frederic Brewster
Paleontology -- Patagonia (Argentina and Chile)
In the Deseado beds, beside _Parastropotherium_, Ameghino has described
_Liarthrus_, based on an upper second premolar and part of another
tooth, but I can see no structural variation from _Parastropotherium_
or indeed from _P. holmbergi_; so I consider this genus as a synonym.
As to the genus _Traspoatherium_, I can not see in it any reason for
making a genus separate from _Parastrapotherium_.
=Parastrapotherium= Ameghino
Parastrapotherium Amegh., 1895, Bol. Inst. Geog. Argen., t. 15, p. 636.
Parastrapotherium Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 449.
Liarthrus Amegh., 1895, Bol. Inst. Geog. Argen., t. 15, p. 641.
Liarthrus Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 451.
Traspoatherium Amegh., 1895, Bol. Inst. Geog. Argen., t. 15, p. 641.
Traspoatherium Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 450.
The genus, in general, is similar to _Astrapotherium_, so that Gaudry
considered it the same, but Ameghino has distinguished it by the tushes
being relatively of smaller size, the lower incisors larger, and by
the presence of pm. 3 in the lower series. The Deseado forms are also
of considerably larger size than the Santa Cruz.
Our material includes a pair of lower jaws, two scapulae, the humerus,
and the lower end of the femur.
No skull has been found in the Deseado. Those from the Santa Cruz are
enormously swollen over the orbits, the massive bone making a skull
wholly unique. The lower jaws are similar to those of _Astrapotherium_,
except that the rami are deeper. The front ends are fused and expanded
making the anterior much enlarged, and causing the incisors to stand at
intervals as in _Coryphodon_. The symphysis is massive and prolonged
backward nearly to premolar 3. The rami are plump and unusually thick.
[Illustration: Fig. 102. Upper dentition of Astrapothericulus
iheringi—½ natural size.]
Of the upper dentition, Ameghino figures only the first molar and
the canine. I have given Ameghino’s figure of the upper dentition of
_Astrapothericulus_, to indicate what this would be like, for the
variation is only slight. The canine is a great tush, not unlike the
incisor-tush of _Pyrotherium_, oval in cross section with the greater
diameter from front to back. The first and second premolars have
disappeared. Premolars 3 and 4 are greatly reduced. The molars are very
like those of _Asmodeus_, large brachydont grinders, composed of an
outer wall, and an anterior and posterior lobe. The external cingulum
is a trace only, and the internal cingulum is developed in varying
degrees. The basin is deep and subdivided by a crista which rises from
the external wall, and as the surface is worn off unites with the
anterior lobe, cutting off a small pit. Behind the posterior lobe is a
small basin, bounded in the rear by a second crista from the rear end
of the external wall, which, as the tooth is worn down, unites with the
posterior lobes, cutting off a small posterior pit, suggestive of that
of homalodontotheres.
Public-domain text, read in full here on John Shaqi.
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