The Deseado Formation of PatagoniaLoomis, Frederic Brewster
Science
The Deseado Formation of Patagonia
Loomis, Frederic Brewster
Paleontology -- Patagonia (Argentina and Chile)
_Parapyrotherium_ is based on a small molar and a tush which Ameghino
first described as _Pyrotherium planum_, later elevating the species
to a genus, designated as _Parapyrotherium_, differentiated by the
transverse crests being low and the valley at either end being blocked
by an intertubercular ridge. Gaudry considered that this genus
represented either the milk teeth of _P. romeri_, or a variation of
that species. I can not see the basis of a new genus in the material.
The genus _Richardoweni_ is based on half of a molar, which has the
transverse crest interrupted in the middle. Too little is known of this
form to base a valid genus or even to associate it with _Pyrotherium_.
_Archaeolophus_ is founded on a small tush and part of an upper molar,
also inadequate material for a genus. It is probably _Pyrotherium_.
_Propyrotherium_ is a smaller form from the Astraponotus beds,
apparently a good genus; the type species being _P. saxeum_, of
considerable smaller size than any of the Deseado species. The
distinctive features of the genus can not be given until more material
is known.
_Carlozitteli_ is based on a small form from the Casamayor with
narrow molars. An incisiform tush is associated with the molar,
which, if correctly associated, would indicate a wide deviation from
_Pyrotherium_, and would probably be an ancestral form. A second
species is reported from the lower Deseado beds, but I am a little
skeptical as to the horizon.
=Pyrotherium= Ameghino
Pyrotherium Amegh., 1889, Actas Acad. Nac. Cienc. Cordoba, t. VI, p.
617.
Pyrotherium Lydekker, 1894, Anal. Mus. La Plata, Palaeontologia
Argentina pt. 3, p. 4.
Pyrotherium Amegh., 1895, Bol. Inst. Geog. Argen., t. 15, p. 609.
Pyrotherium Amegh., 1897, Bol. Inst. Geog. Argen., t. 18, p. 441.
Pyrotherium Amegh., 1902, Anal. Mus. Nac. Buenos Aires, ser. 3, t. 1,
p. 19-43.
Pyrotherium Amegh., 1902, Anal. Mus. Nac. Buenos Aires, ser. 3, t. 1,
p. 223-4.
Pyrotherium Gaudry, 1909, Anal. Palaeontologie, t. 4, p. 1-28.
Parapyrotherium Amegh., 1902, Anal. Mus. Nac. Buenos Aires, ser. 3, t.
1, p. 29.
The type species of the genus is _P. romeri_, which is however a rare
species, most of the material and the best known belonging to _P.
sorondoi_. The Amherst Collection contains a skull, complete except
that the top of the brain case is crushed in and the parietals lost;
a second skull with the full upper dentition but lacking the cranium;
four lower jaws; two isolated tushes; the atlas, axis, and cervicals
3 and 4; the humerus; the proximal end of the femur; and part of the
front foot; all from the Chico del Chubut west of Puerto Visser. Gaudry
had upper and lower dentition and the fore and hind limbs except the
feet. Ameghino described the upper and lower dentitions and a fore
foot, so that with our material we now have a basis for a fairly
complete discussion, the vertebral column being the major part which is
still lacking.
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