As any variation in the structure of the reproductive organs, combined with changed function, is a rare event, the following cases are worth giving in detail. My attention was first called to the subject by observing, in 1862, a long-styled plant, descended from a self-fertilised long-styled parent, which had some of its flowers in an anomalous state, namely, with the stamens placed low down in the corolla as in the ordinary long-styled form, but with the pistils so short that the stigmas stood on a level with the anthers. These stigmas were nearly as globular and as smooth as in the short-styled form, instead of being elongated and rough as in the long-styled form. Here, then, we have combined in the same flower, the short stamens of the long-styled form with a pistil closely resembling that of the short-styled form. But the structure varied much even on the same umbel: for in two flowers the pistil was intermediate in length between that of the long and that of the short-styled form, with the stigma elongated as in the former, and smooth as in the latter; and in three other flowers the structure was in all respects like that of the long-styled form. These modifications appeared to me so remarkable that I fertilised eight of the flowers with their own pollen, and obtained five capsules, which contained on an average 43 seeds; and this number shows that the flowers had become abnormally fertile in comparison with those of ordinary long- styled plants when self-fertilised. I was thus led to examine the plants in several small collections, and the result showed that the equal-styled variety was not rare. TABLE 5.31. Primula Sinensis. Preponderance of long-styled over the short-styled form. Column 1: Name of owner or place. Column 2: Long-styled form. Column 3: Short-styled form. Column 4: equal-styled variety. Mr. Horwood : 0 : 0 : 17. Mr. Duck : 20 : 0 : 9. Baston : 30 : 18 : 15. Chichester : 12 : 9 : 2. Holwood : 42 : 12 : 0. High Elms : 16 : 0 : 0. Westerham : 1 : 5 : 0. My own plants from purchased seeds : 13 : 7 : 0. Total : 134 : 51 : 43.