projected above the anthers, but neither were properly long-styled or equal- styled; and in the one in which the stigma projected most, the grains were in diameter to those in the other plant, in which the stigma projected less, as 100 to 88; whereas the difference between the grains from perfectly characterised long-styled and short-styled plants is as 100 to 57. So that these two plants were in an intermediate condition. To return to the 17 plants in the first line of Table 5.31: from the relative position of their stigmas and anthers, they could hardly fail to fertilise themselves; and accordingly four of them spontaneously yielded no less than 180 capsules; of these Mr. Horwood selected eight fine capsules for sowing; and they included on an average 54.8 seeds, with a maximum of 72. He gave me thirty other capsules, taken by hazard, of which twenty-seven contained good seeds, averaging 35.5, with a maximum of 70; but if six poor capsules, each with less than 13 seeds, be excluded, the average rises to 42.5. These are higher numbers than could be expected from either well- characterised form if self-fertilised; and this high degree of fertility accords with the view that the male organs belonged to one form, and the female organs partially to the other form; so that a self-union in the case of the equal- styled variety is in fact a legitimate union. The seed saved from the above seventeen self-fertilised equal-styled plants produced sixteen plants, which all proved equal-styled, and resembled their parents in all the above-specified respects. The stamens, however, in one plant were seated higher up the tube of the corolla than in the true long-styled form; in another plant almost all the anthers were contabescent. These sixteen plants were the grandchildren of the four original plants, which it is believed were equal-styled; so that this abnormal condition was faithfully transmitted, probably through three, and certainly through two generations. The fertility of one of these grandchildren was carefully observed: six flowers were fertilised with pollen from the same flower, and produced six capsules, containing on an average 68 seeds, with a maximum of 82, and a minimum of 40. Thirteen capsules spontaneously self-fertilised yielded an average of 53.2 seeds, with the astonishing maximum in one of 97 seeds. In no legitimate union has so high an average as 68 seeds been observed by me, or nearly so high a maximum as 82 and 97. These plants, therefore, not only have lost their proper heterostyled structure and peculiar functional powers, but have acquired an abnormal grade of fertility--unless, indeed, their high fertility may be accounted for by the stigmas receiving pollen from the circumjacent anthers at exactly the most favourable period.