The Doctrine of Evolution: Its Basis and Its ScopeCrampton, Henry Edward
Science
The Doctrine of Evolution: Its Basis and Its Scope
Crampton, Henry Edward
Evolution
But that is a metaphysical matter, with which we need not concern
ourselves in this discussion; the important point is that among the
everyday happenings of life are processes that are quite competent to
account for the condition of adaptation exhibited by various animal forms.
These processes are real and natural, not imaginative or artificial, and
so they will remain even though it will become clear that much is still to
be learned about the causes of variation and the course of biological
inheritance. Darwin was the first to contend that natural selection is but
a part of nature's method of accomplishing evolution. As such it is
content to recognize variations and does not concern itself with the
origin of modifications; it accepts the obvious fact that congenital
variations are inherited, although it leaves the question as to how they
are inherited for further examination. Because the doctrine of natural
selection does not profess to answer all the questions propounded by
scientific inquisitiveness, it must not be supposed that it fails in its
immediate purpose of giving a natural explanation of how evolution may be
partly accounted for.
* * * * *
Before proceeding to the post-Darwinian investigations that have done so
much to amplify the account of natural evolution, let us consider the
contrasted explanation given by Lamarck and his followers. As we have
stated earlier, Lamarckianism is the name given to the doctrine that
modifications other than those due to congenital factors may enter into
the heritage of a species, and may add themselves to those already
combined as the peculiar characteristics of a particular species. Let us
take the giraffe and its long neck as a concrete example. The great length
of this part is obviously an adaptive character, enabling the animal to
browse upon the softer leafy shoots of shrubs and trees. The vertebral
column of the neck comprises just the same number of bones that are
present in the short-necked relatives of this form, so that we are
justified in accepting as a fact the evolution of the giraffe's long neck
by the lengthening of each one of originally shorter vertebrae. The
Lamarckian explanation of this fact would be that the earliest forms in
the ancestry of the giraffe as such stretched their necks as they fed, and
that this peculiar function with its correlated structural modification
became habitual. The slight increase brought about by any single
individual would be inherited and transmitted to the giraffes of the next
generation; in other words, an individually acquired character would be
inherited. The young giraffes of this next generation would then begin,
not where their parents did, but from an advanced condition. Thus, by
continued stretching of the neck and by continued transmission of the
elongated condition, the great length of this part of the body in the
modern giraffe would be attained.
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