The 'superior maxillaries' here swell into importance, and constitute
the whole of the outer wall, and, sometimes, a part of the roof. The
jaws are the weapons of offence and defence; and as much space as
possible is devoted to the insertion of those muscles that will enable
the animal to seize and to hold his prey. One of the most powerful of
them, the 'masseter', rises from the superior maxillary bone, and
spreads over its whole extent: therefore, that bone is developed, while
the nasal bone is compressed into a very small space. The substitution
of a portion of cartilage, instead of bone, at the posterior part of the
orbital ring, in order to give more play for the coracoid process of the
posterior maxillary, round which the temporal bone is wrapped, is a
contrivance of the same nature.
The scent of the dog is not sacrificed or impaired by the apparent
diminution of the nasals; for the cavity enlarges considerably upward,
and is occupied chiefly by the 'ethmoid bone', which, having the greater
portion of nervous pulp spread on it, seems to have most to do with the
sense of smell.
The nasal bones of the dog are essentially different from those of the
horse, cattle, and sheep. They commence, indeed, as high up in the face
as those of the horse, their superior extremities being opposite to the
lachrymal gland; but that commencement is an apex or point varying
materially in different breeds. They form, altogether, one sharp
projection, and are received within breeds these processes extend nearly
one-third of the length of the nasals.
The superior maxillary (3.3.) takes the situation of the nasal (2.),
pushes the lachrymal bone (4.) out of its place, and almost annihilates
it, reaches the frontal bone (7.) and expands upon it, and forms with it
the same denticulated suture which is to be seen in the nasal. The
action of the muscle between these bones, and for the development of
which all this sacrifice is made, is exceedingly powerful. The strength
of this muscle in a large dog is almost incredible: the sutures between
these bones must possess corresponding strength; and so strong is the
union between them, that, in many old dogs, the suture between the
superior maxillary and frontal bones is nearly obliterated, and that
between the nasal and frontal maxillary quite effaced.
Public-domain text, read in full here on John Shaqi.
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