Often the organic substance has neither been preserved nor replaced,
but the _form_ has been retained by means of molds and casts.
Permanent impressions, or molds, may be made in sediments not only by
the hard parts of organisms, but also by such soft and perishable
parts as the leaves of plants, and, in the rarest instances, by the
skin of animals and the feathers of birds. In fine-grained limestones
even the imprints of jellyfish have been retained.
The different kinds of molds and casts may be illustrated by means of
a clam shell and some moist clay, the latter representing the
sediments in which the remains of animals and plants are entombed.
Imbedding the shell in the clay and allowing the clay to harden, we
have a _mold of the exterior_ of the shell, as is seen on cutting the
clay matrix in two and removing the shell from it. Filling this mold
with clay of different color, we obtain a _cast of the exterior_,
which represents accurately the original form and surface markings of
the shell. In nature, shells and other relics of animals or plants are
often removed by being dissolved by percolating waters, and the molds
are either filled with sediments or with minerals deposited from
solution.
Where the fossil is hollow, a _cast of the interior_ is made in the
same way. Interior casts of shells reproduce any markings on the
inside of the valves, and casts of the interior of the skulls of
ancient vertebrates show the form and size of their brains.
=Imperfection of the life record.= At the present time only the
smallest fraction of the life on earth ever gets entombed in rocks now
forming. In the forest great fallen tree trunks, as well as dead
leaves, decay, and only add a little to the layer of dark vegetable
mold from which they grew. The bones of land animals are, for the most
part, left unburied on the surface and are soon destroyed by chemical
agencies. Even where, as in the swamps of river, flood plains and in
other bogs, there are preserved the remains of plants, and sometimes
insects, together with the bones of some animal drowned or mired, in
most cases these swamp and bog deposits are sooner or later destroyed
by the shifting channels of the stream or by the general erosion of
the land.
In the sea the conditions for preservation are more favorable than on
land; yet even here the proportion of animals and plants whose hard
parts are fossilized is very small compared with those which either
totally decay before they are buried in slowly accumulating sediments
or are ground to powder by waves and currents.
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