In many cases the extinction of a type has been gradual; in other instances apparently abrupt. There is no evidence that any organism once become extinct has ever reappeared. The duration of a species in time, or its "vertical range" through the strata, varies greatly. Some species are limited to a stratum a few feet in thickness; some may range through an entire formation and be found but little modified in still higher beds. A formation may thus often be divided into zones, each characterized by its own peculiar species. As a rule, the simpler organisms have a longer duration as species, though not as individuals, than the more complex. 3. _The larger zoölogical and botanical groupings survive longer than the smaller._ Species are so short-lived that a single geological epoch may be marked by several more or less complete extinctions of the species of its fauna-flora and their replacement by other species. A genus continues with new species after all the species with which it began have become extinct. Families survive genera, and orders families. Classes are so long-lived that most of those which are known from the earliest formations are represented by living forms, and no subkingdom has ever become extinct. Thus, to take an example from the stony corals,--the _zoantharia_,--the particular characters--which constituted a certain _species_--_Facosites niagarensis_--of the order are confined to the Niagara series. Its _generic_ characters appeared in other species earlier in the Silurian and continued through the Devonian. Its _family_ characters, represented in different genera and species, range from the Ordovician to the close of the Paleozoic; while the characters which it shares with all its order, the Zoantharia, began in the Cambrian and are found in living species. 4. _The change in organisms has been gradual._ The fossils of each life zone and of each formation of a conformable series closely resemble, with some explainable exceptions, those of the beds immediately above and below. The animals and plants which tenanted the earth during any geological epoch are so closely related to those of the preceding and the succeeding epochs that we may consider them to be the descendants of the one and the ancestors of the other, thus accounting for the resemblance by heredity. It is therefore believed that the species of animals and plants now living on the earth are the descendants of the species whose remains we find entombed in the rocks, and that the chain of life has been unbroken since its beginning.