Such lowly forms were destitute of any hard parts and could leave no
evidence of their existence in the record of the rocks. And of their
supposed descendants we find so few traces in the pre-Cambrian strata
that the first steps in organic evolution must be supplied from such
analogies in embryology as the following. The fertilized ovum, the
cell with which each animal begins its life, grows and multiplies by
cell division, and develops into a hollow globe of cells called the
_blastosphere_. This stage is succeeded by the stage of the
_gastrula_,--an ovoid or cup-shaped body with a double wall of cells
inclosing a body cavity, and with an opening, the primitive mouth.
Each of these early embryological stages is represented by living
animals,--the undivided cell by the _protozoa_, the blastosphere by
some rare forms, and the gastrula in the essential structure of the
_coelenterates_,--the subkingdom to which the fresh-water hydra and
the corals belong. All forms of animal life, from the coelenterates to
the mammals, follow the same path in their embryological development
as far as the gastrula stage, but here their paths widely diverge,
those of each subkingdom going their own separate ways.
We may infer, therefore, that during the pre-Cambrian periods organic
evolution followed the lines thus dimly traced. The earliest
one-celled protozoa were probably succeeded by many-celled animals of
the type of the blastosphere, and these by gastrula-like organisms.
From the gastrula type the higher subdivisions of animal life
probably diverged, as separate branches from a common trunk. Much or
all of this vast differentiation was accomplished before the opening
of the next era; for all the subkingdoms are represented in the
Cambrian except the vertebrates.
=Evidences of pre-Cambrian life.= An indirect evidence of life during
the pre-Cambrian periods is found in the abundant and varied fauna of
the next period; for, if the theory of evolution is correct, the
differentiation of the Cambrian fauna was a long process which might
well have required for its accomplishment a large part of pre-Cambrian
time.
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