During the long ages of the Cambrian the trilobite varied greatly.
Again and again new species and genera appeared, while the older types
became extinct. For this reason and because of their abundance,
trilobites are used in the classification of the Cambrian system. The
Lower Cambrian is characterized by the presence of a trilobitic fauna
in which the genus Olenellus is predominant. This, the _Olenellus
Zone_, is one of the most important platforms in the entire geological
series; for, the world over, it marks the beginning of Paleozoic time,
while all underlying strata are classified as pre-Cambrian. The Middle
Cambrian is marked by the genus Paradoxides, and the Upper Cambrian by
the genus Olenus. Some of the Cambrian trilobites were giants,
measuring as much as two feet long, while others were the smallest of
their kind, a fraction of an inch in length.
Another type of crustacean which lived in the Cambrian and whose order
is still living is illustrated in Figure 269.
[Illustration: Fig. 269. A Phyllopod]
=Worms.= Trails and burrows of worms have been left on the sea beaches
and mud flats of all geological times from the Algonkian to the
present.
=Brachiopods.= These soft-bodied animals, with bivalve shells and two
interior armlike processes which served for breathing, appeared in the
Algonkian, and had now become very abundant. The two valves of the
brachiopod shell are unequal in size, and in each valve a line drawn
from the beak to the base divides the valve into two equal parts
(Fig. 270). It may thus be told from the pelecypod mollusk, such as the
clam, whose two valves are not far from equal in size, each being
divided into unequal parts by a line dropped from the beak (Fig.272).
[Illustration: Fig. 270. A Cambrian Articulate Brachiopod, Orthis]
[Illustration: Fig. 271. Cambrian Inarticulate Brachiopods
A, Lingulella; B, Discina]
Brachiopods include two orders. In the most primitive order--that of
the _inarticulate_ brachiopods--the two valves are held together only
by muscles of the animal, and the shell is horny or is composed of
phosphate of lime. The _Discina_, which began in the Algonkian, is of
this type, as is also the _Lingulella_ of the Cambrian (Fig. 271). Both
of these genera have lived on during the millions of years of geological
time since their introduction, handing down from generation to
generation with hardly any change to their descendants now living off
our shores the characters impressed upon them at the beginning.
The more highly organized _articulate_ brachiopods have valves of
carbonate of lime more securely joined by a hinge with teeth and
sockets (Fig. 270). In the Cambrian the inarticulates predominate,
though the articulates grow common toward the end of the period.
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