=Teleosts.= The great group of fishes called the teleosts, or those
with complete bony skeletons, to which most modern fishes belong, may
be mentioned here, although in the Devonian they had not yet appeared.
The teleosts are a highly specialized type, adapted most perfectly to
their aquatic environment. Heavy armor has been discarded, and
reliance is placed instead on swiftness. The skeleton is completely
ossified and the notochord removed. The vertebræ have been
economically withdrawn from the tail, and the cartilaginous axis of
the side fins has been found unnecessary. The air sac has become a
swim bladder. In this complete specialization they have long since
lost the possibility of evolving into higher types.
It is interesting to note that the modern teleosts in their
embryological growth pass through the stages which characterized the
maturity of their Devonian ancestors; their skeleton is cartilaginous
and their tail fin vertebrated.
CHAPTER XIX
THE CARBONIFEROUS
The Carboniferous system is so named from the large amount of coal
which it contains. Other systems, from the Devonian on, are coal
bearing also, but none so richly and to so wide an extent. Never
before or since have the peculiar conditions been so favorable for the
formation of extensive coal deposits.
With few exceptions the Carboniferous strata rest on those of the
Devonian without any marked unconformity; the one period passed
quietly into the other, with no great physical disturbances.
The Carboniferous includes three distinct series. The lower is called
the _Mississippian_, from the outcrop of its formations along the
Mississippi River in central and southern Illinois and the adjacent
portions of Iowa and Missouri. The middle series is called the
_Pennsylvanian_ (or Coal Measures), from its wide occurrence over
Pennsylvania. The upper series is named the _Permian_, from the
province of Perm in Russia.
=The Mississippian series.= In the interior the Mississippian is
composed chiefly of limestones, with some shales, which tell of a
clear, warm, epicontinental sea swarming with crinoids, corals, and
shells, and occasionally clouded with silt from the land.
In the eastern region, New York had been added by uplift to the
Appalachian land which now was united to the northern area. From
eastern Pennsylvania southward there were laid in a subsiding trough,
first, thick sandstones (the Pocono sandstone), and later still
heavier shales,--the two together reaching the thickness of four
thousand feet and more. We infer a renewed uplift of Appalachia
similar to that of the later epochs of the Devonian, but as much less
in amount as the volume of sediments is smaller.
The Pennsylvanian Series
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