=Fishes= still held to the Devonian types, with the exception that the
strange ostracoderms now had perished.
=Amphibians.= The vertebrates had now followed the arthropods and the
mollusks upon the land, and developed a higher type adapted to the new
environment. Amphibians--the class to which frogs and salamanders
belong--now appear, with lungs for breathing air and with limbs for
locomotion on the land. Most of the Carboniferous amphibians were shaped
like the salamander, with weak limbs adapted more for crawling than for
carrying the body well above the ground. some legless, degenerate forms
were snakelike in shape.
[Illustration: Fig. 310. A Carboniferous Dragon Fly
One tenth natural size]
The earliest amphibians differ from those of to-day in a number of
respects. They were connecting types linking together fishes, from
which they were descended, with reptiles, of which they were the
ancestors. They retained the evidence of their close relationship with
the Devonian fishes in their cold blood, their gills and aquatic habit
during their larval stage, their teeth with dentine infolded like
those of the Devonian ganoids but still more intricately, and their
biconcave vertebræ which never completely ossified. These, the
highest vertebrates of the time, had not yet advanced beyond the
embryonic stage of the more or less cartilaginous skeleton and the
persistent notochord.
[Illustration: Fig. 311. A Carboniferous Amphibian]
[Illustration: Fig. 312. Transverse Section of
Segment of Tooth of Carboniferous Amphibian]
On the other hand, the skull of the Carboniferous amphibians was made
of close-set bony plates, like the skull of the reptile, rather than
like that of the frog, with its open spaces (Figs. 313 and 314).
Unlike modern amphibians, with their slimy skin, the Carboniferous
amphibians wore an armor of bony scales over the ventral surface and
sometimes over the back as well.
[Illustration: Fig. 313. Skull of a Permian Amphibian from Texas]
[Illustration: Fig. 314. Skull of a Frog]
It is interesting to notice from the footprints and skeletons of these
earliest-known vertebrates of the land what was the primitive number
of digits. The Carboniferous amphibians had five-toed feet, the
primitive type of foot, from which their descendants of higher orders,
with a smaller number of digits, have diverged.
The Carboniferous was the age of lycopods and amphibians, as the
Devonian had been the age of rhizocarps and fishes.
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