The lowest in grade and nearest, perhaps, to the ancestral type of
vertebrates, was the problematic creature, an inch or so long, of
Figure 297. Note the circular mouth not supplied with jaws, the
lack of paired fins, and the symmetric tail fin, with the column
of cartilaginous, ringlike vertebrae running through it to the
end. The animal is probably to be placed with the jawless lampreys
and hags,--a group too low to be included among true fishes.
OSTRACODERMS. This archaic group, long since extinct, is also too
lowly to rank among the true fishes, for its members have neither
jaws nor paired fins. These small, fishlike forms were cased in
front with bony plates developed in the skin and covered in the
rear with scales. The vertebrae were not ossified, for no trace of
them has been found.
DEVONIAN FISHES. The TRUE FISHES of the Devonian can best be
understood by reference to their descendants now living. Modern
fishes are divided into several groups: SHARKS and their allies;
DIPNOANS; GANOIDS, such as the sturgeon and gar; and TELEOSTS,--
most common fishes, such as the perch and cod.
SHARKS. Of all groups of living fishes the sharks are the oldest
and still retain most fully the embryonic characters of their
Paleozoic ancestors. Such characters are the cartilaginous
skeleton, and the separate gill slits with which the throat wall
is pierced and which are arranged in line like the gill openings
of the lamprey. The sharks of the Silurian and Devonian are known
to us chiefly by their teeth and fin spines, for they were
unprotected by scales or plates, and were devoid of a bony
skeleton. Figure 299 is a restoration of an archaic shark from a
somewhat higher horizon. Note the seven gill slits and the
lappetlike paired fins. These fins seem to be remnants of the
continuous fold of skin which, as embryology teaches, passed from
fore to aft down each side of the primitive vertebrate.
Devonian sharks were comparatively small. They had not evolved
into the ferocious monsters which were later to be masters of the
seas.
DIPNOANS, OR LUNG FISHES. These are represented to-day by a
few peculiar fishes and are distinguished by some high structures
which ally them with amphibians. An air sac with cellular spaces
is connected with the gullet and serves as a rudimentary lung. It
corresponds with the swim bladder of most modern fishes, and
appears to have had a common origin with it. We may conceive that
the primordial fishes not only had gills used in breathing air
dissolved in water, but also developed a saclike pouch off the
gullet. This sac evolved along two distinct lines. On the line of
the ancestry of most modern fishes its duct was closed and it
became the swim bladder used in flotation and balancing. On
another line of descent it was left open, air was swallowed into
it, and it developed into the rudimentary lung of the dipnoans and
into the more perfect lungs of the amphibians and other air-
breathing vertebrates.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account