Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The wide comparative studies of the scientists I have named have
further shown that in the case of the discoblastic higher vertebrates
(the three classes of amniotes) the primitive mouth of the embryonic
disc, which was long looked for in vain, is found always, and is
nothing else than the familiar “primitive groove.” Of this we shall see
more as we proceed. Meantime we realise that gastrulation may be
reduced to one and the same process in all the vertebrates. Moreover,
the various forms it takes in the invertebrates can always be reduced
to one of the four types of segmentation described above. In relation
to the distinction between total and partial segmentation, the grouping
of the various forms is as follows:—
I. Palingenetic
(primitive) segmentation. 1. Equal segmentation
(bell-gastrula). A. Total segmentation (without independent
food-yelk). II. Cenogenetic segmentation
(modified by adaptation). 2. Unequal segmentation (hooded
gastrula). 3. Discoid segmentation (discoid gastrula). B.
Partial segmentation (with independent food-yelk). 4. Superficial
segmentation
(spherical gastrula).
The lowest metazoa we know—namely, the lower zoophyta (sponges, simple
polyps, etc.)—remain throughout life at a stage of development which
differs little from the gastrula; their whole body consists of two
layers of cells. This is a fact of extreme importance. We see that man,
and also other vertebrates, pass quickly through a stage of development
in which they consist of two layers, just as these lower zoophyta do
throughout life. If we apply our biogenetic law to the matter, we at
once reach this important conclusion. “Man and all the other animals
which pass through the two-layer stage, or gastrula-form, in the course
of their embryonic development, must descend from a primitive simple
stem-form, the whole body of which consisted throughout life (as is the
case with the lower zoophyta to-day) merely of two cell-strata or
germinal layers.” We will call this primitive stem-form, with which we
shall deal more fully later on, the _ gastræa_—that is to say,
“primitive-gut animal.”
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