Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
Now a second, narrower but longer, cavity arises by a process of
folding at the lower pole, and by the falling away from each other of
the white entoderm-cells (Figs. 41–44 _N_). This is the primitive
gut-cavity or the gastric cavity of the gastrula, progaster or
archenteron. It was first observed in the ovum of the amphibia by
Rusconi, and so called the Rusconian cavity. The reason of its peculiar
narrowness here is that it is, for the most part, full of yelk-cells of
the entoderm. These also stop up the whole of the wide opening of the
primitive mouth, and form what is known as the “yelk-stopper,” which is
seen freely at the white round spot at the south pole (_P_). Around it
the ectoderm is much thicker, and forms the border of the primitive
mouth, the most important part of the embryo (Fig. 44 _k, k′_). Soon
the primitive gut-cavity stretches further and further at the expense
of the segmentation-cavity (_F_), until at last the latter disappears
altogether. The two cavities are only separated by a thin partition
(Fig. 43 _s_). With the formation of the primitive gut our frog-embryo
has reached the gastrula stage, though it is clear that this
cenogenetic amphibian gastrula is very different from the real
palingenetic gastrula we have considered (Figs. 30–36).
In the growth of this hooded gastrula we cannot sharply mark off the
various stages which we distinguish successively in the bell-gastrula
as morula and gastrula. Nevertheless, it is not difficult to reduce the
whole cenogenetic or disturbed development of this amphigastrula to the
true palingenetic formation of the archigastrula of the amphioxus.
Fig.46. Embryonic vesicle of triton. Fig. 46—Embryonic vesicle of
triton (_blastula_), outer view, with the transverse fold of the
primitive mouth (_u_). (From _Hertwig._)
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