Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The bird’s egg is particularly important for our purpose, because most
of the chief studies of the development of the vertebrates are based on
observations of the hen’s egg during hatching. The mammal ovum is much
more difficult to obtain and study, and for this practical and obvious
reason very rarely thoroughly investigated. But we can get hens’ eggs
in any quantity at any time, and, by means of artificial incubation,
follow the development of the embryo step by step. The bird’s egg
differs considerably from the tiny mammal ovum in size, a large
quantity of food-yelk accumulating within the original yelk or the
protoplasm of the ovum. This is the yellow ball which we commonly call
the yolk of the egg. In order to understand the bird’s egg aright—for
it is very often quite wrongly explained—we must examine it in its
original condition, and follow it from the very beginning of its
development in the bird’s ovary. We then see that the original ovum is
a quite small, naked, and simple cell with a nucleus, not differing in
either size or shape from the original ovum of the mammals and other
animals (cf. Fig. 13 _ E_). As in the case of all the craniota (animals
with a skull), the original or primitive ovum (_protovum_) is covered
with a continuous layer of small cells. This membrane is the follicle,
from which the ovum afterwards issues. Immediately underneath it the
structureless yelk-membrane is secreted from the yelk.
The small primitive ovum of the bird begins very early to take up into
itself a quantity of food-stuff through the yelk-membrane, and work it
up into the “yellow yelk.” In this way the ovum
enters on its second stage (the metovum), which is many times larger
than the first, but still only a single enlarged cell. Through the
accumulation of the store of yellow yelk within the ball of protoplasm
the nucleus it contains (the germinal vesicle) is forced to the surface
of the ball. Here it is surrounded by a small quantity of protoplasm,
and with this forms the lens-shaped formative yelk (Fig. 15 _b_). This
is seen on the yellow yelk-ball, at a certain point of the surface, as
a small round white spot—the “tread” (_cicatricula_). From this point a
thread-like column of white nutritive yelk (_d_), which contains no
yellow yelk-granules, and is softer than the yellow food-yelk, proceeds
to the middle of the yellow yelk-ball, and forms there a small central
globule of white yelk (Fig. 15 _ d_). The whole of this white yelk is
not sharply separated from the yellow yelk, which shows a slight trace
of concentric layers in the hard-boiled egg (Fig. 15 _c_). We also find
in the hen’s egg, when we break the shell and take out the yelk, a
round small white disk at its surface which corresponds to the tread.
But this small white “germinal disk” is now further developed, and is
really the gastrula of the chick. The body of the chick is formed from
it alone. The whole white and yellow yelk-mass is without any
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