Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The impregnated ovum of this coral (Figure 1.29 A, B) first splits
into two equal cells (C). First, the nucleus of the stem-cell and its
central body divide into two halves. These recede from and repel each
other, and act as centres of attraction on the surrounding protoplasm;
in consequence of this, the protoplasm is constricted by a circular
furrow, and, in turn, divides into two halves. Each of the two
segmentation-cells thus produced splits in the same way into two equal
cells. The four segmentation-cells (grand-daughters of the stem-cell)
lie in one plane. Now, however, each of them subdivides into two equal
halves, the cleavage of the nucleus again preceding that of the
surrounding protoplasm. The eight cells which thus arise break into
sixteen, these into thirty-two, and then (each being constantly
halved) into sixty-four, 128, and so on.* (* The number of
segmentation-cells thus produced increases geometrically in the
original gastrulation, or the purest palingenetic form of cleavage.
However, in different animals the number reaches a different height,
so that the morula, and also the blastula, may consist sometimes of
thirty-two, sometimes of sixty-four, and sometimes of 128, or more,
cells.) The final result of this repeated cleavage is the formation of
a globular cluster of similar segmentation-cells, which we call the
mulberry-formation or morula. The cells are thickly pressed together
like the parts of a mulberry or blackberry, and this gives a lumpy
appearance to the surface of the sphere (Figure E).* (* The
segmentation-cells which make up the morula after the close of the
palingenetic cleavage seem usually to be quite similar, and to present
no differences as to size, form, and composition. That, however, does
not prevent them from differentiating into animal and vegetative
cells, even during the cleavage.)
When the cleavage is thus ended, the mulberry-like mass changes into a
hollow globular sphere. Watery fluid or jelly gathers inside the
globule; the segmentation-cells are loosened, and all rise to the
surface. There they are flattened by mutual pressure, and assume the
shape of truncated pyramids, and arrange themselves side by side in
one regular layer (Figures F, G). This layer of cells is called the
germinal membrane (or blastoderm); the homogeneous cells which compose
its simple structure are called blastodermic cells; and the whole
hollow sphere, the walls of which are made of the preceding, is called
the blastula or blastosphere.* (* The blastula of the lower animals
must not be confused with the very different blastula of the mammal,
which is properly called the gastrocystis or blastocystis. This
cenogenetic gastrocystis and the palingenetic blastula are sometimes
very wrongly comprised under the common name of blastula or vesicula
blastodermica.)
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