Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The wide comparative studies of the scientists I have named have
further shown that in the case of the discoblastic higher vertebrates
(the three classes of amniotes) the primitive mouth of the embryonic
disc, which was long looked for in vain, is found always, and is
nothing else than the familiar "primitive groove." Of this we shall
see more as we proceed. Meantime we realise that gastrulation may be
reduced to one and the same process in all the vertebrates. Moreover,
the various forms it takes in the invertebrates can always be reduced
to one of the four types of segmentation described above. In relation
to the distinction between total and partial segmentation, the
grouping of the various forms is as follows:--
1. Palingenetic (primitive segmentation)
1.1. Equal segmentation (bell-gastrula).
1.1.A. Total segmentation (without independent food-yelk).
2. Cenogenetic segmentation (modified by adaptation).
2.2. Unequal segmentation (hooded gastrula).
2.2.A. Total segmentation (without independent food-yelk).
2.3. Discoid segmentation (discoid gastrula).
2.3.B. Partial segmentation (with independent food-yelk).
2.4. Superficial segmentation (spherical gastrula).
2.4.B. Partial segmentation (with independent food-yelk).
The lowest metazoa we know--namely, the lower zoophyta (sponges,
simple polyps, etc.)--remain throughout life at a stage of development
which differs little from the gastrula; their whole body consists of
two layers of cells. This is a fact of extreme importance. We see that
man, and also other vertebrates, pass quickly through a stage of
development in which they consist of two layers, just as these lower
zoophyta do throughout life. If we apply our biogenetic law to the
matter, we at once reach this important conclusion. "Man and all the
other animals which pass through the two-layer stage, or
gastrula-form, in the course of their embryonic development, must
descend from a primitive simple stem-form, the whole body of which
consisted throughout life (as is the case with the lower zoophyta
to-day) merely of two cell-strata or germinal layers." We will call
this primitive stem-form, with which we shall deal more fully later
on, the gastraea--that is to say, "primitive-gut animal."
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