Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
This reduction is tolerably easy and confident in the case of the
small ovum of our deep-sea bony fish, but it becomes difficult and
uncertain in the case of the large ova that we find in the majority of
the other fishes and in all the reptiles and birds. In these cases the
food-yelk is, in the first place, comparatively colossal, the
formative yelk being almost invisible beside it; and, in the second
place, the food-yelk contains a quantity of different elements, which
are known as "yelk-granules, yelk-globules, yelk-plates, yelk-flakes,
yelk-vesicles," and so on. Frequently these definite elements in the
yelk have been described as real cells, and it has been wrongly stated
that a portion of the embryonic body is built up from these cells.
This is by no means the case. In every case, however large it is--and
even when cell-nuclei travel into it during the cleavage of the
border--the nutritive yelk remains a dead accumulation of food, which
is taken into the gut during embryonic development and consumed by the
embryo. The latter develops solely from the living formative yelk of
the stem-cell. This is equally true of the ova of our small bony
fishes and of the colossal ova of the primitive fishes, reptiles, and
birds.
(FIGURE 1.54. Discoid gastrula (discogastrula) of a bony fish. e
ectoderm, i entoderm, w border-swelling or primitive mouth, n
albuminous globule of the nutritive yelk, f fat-globule of same, c
external membrane (ovolemma), d partition between entoderm and
ectoderm (earlier the segmentation-cavity).)
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