Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
How we are to explain these phylogenetic relations in the present
state of our knowledge, and what place is assigned to the vertebrates
in the animal ancestral tree, will be considered later (Chapter 2.20).
For the present our task is to make plainer the vertebrate character
of man, and especially to point out the chief peculiarities of
organisation by which the vertebrate stem is profoundly separated from
the other eleven stems of the animal kingdom. Only after these
comparative-anatomical considerations shall we be in a position to
attack the difficult question of our embryology. The development of
even the simplest and lowest vertebrate from the simple chordula
(Figures 1.83 to 1.86) is so complicated and difficult to follow that
it is necessary to understand the organic features of the fully-formed
vertebrate in order to grasp the course of its embryonic evolution.
But it is equally necessary to confine our attention, in this general
anatomic description of the vertebrate-body, to the essential facts,
and pass by all the unessential. Hence, in giving now an ideal
anatomic description of the chief features of the vertebrate and its
internal organisation, I omit all the subordinate points, and restrict
myself to the most important characteristics.
Much, of course, will seem to the reader to be essential that is only
of subordinate and secondary interest, or even not essential at all,
in the light of comparative anatomy and embryology. For instance, the
skull and vertebral column and the extremities are non-essential in
this sense. It is true that these parts are very important
PHYSIOLOGICALLY; but for the MORPHOLOGICAL conception of the
vertebrate they are not essential, because they are only found in the
higher, not the lower, vertebrates. The lowest vertebrates have
neither skull nor vertebrae, and no extremities or limbs. Even the
human embryo passes through a stage in which it has no skull or
vertebrae; the trunk is quite simple, and there is yet no trace of
arms and legs. At this stage of development man, like every other
higher vertebrate, is essentially similar to the simplest vertebrate
form, which we now find in only one living specimen. This one lowest
vertebrate that merits the closest study--undoubtedly the most
interesting of all the vertebrates after man--is the famous lancelet
or amphioxus, to which we have already often referred. As we are going
to study it more closely later on (Chapters 2.16 and 2.17), I will
only make one or two passing observations on it here.
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