The kidneys, which act as organs of excretion or urinary organs in all vertebrates, have a very different and elaborate construction in the various sections of this stem; we will consider them further in Chapter 2.29. Here I need only mention that in our hypothetical primitive vertebrate they probably had the same form as in the actual amphioxus--the primitive kidneys (protonephra). These are originally made up of a double row of little canals, which directly convey the used-up juices or the urine out of the body-cavity (Figure 1.102 n). The inner aperture of these pronephridial canals opens with a ciliated funnel into the body-cavity; the external aperture opens in lateral grooves of the epidermis, a couple of longitudinal grooves in the lateral surface of the outer skin (Figure 1.102 b). The pronephridial duct is formed by the closing of this groove to the right and left at the sides. In all the craniota it develops at an early stage in the horny plate; in the amphioxus it seems to be converted into a wide cavity, the atrium, or peribranchial space. Next to the kidneys we have the sexual organs of the vertebrate. In most of the members of this stem the two are united in a single urogenital system; it is only in a few groups that the urinary and sexual organs are separated (in the amphioxus, the cyclostoma, and some sections of the fish-class). In man and all the higher vertebrates the sexual apparatus is made up of various parts, which we will consider in Chapter 2.29. But in the two lowest classes of our stem, the acrania and cyclostoma, they consist merely of simple sexual glands or gonads, the ovaries of the female sex and the testicles (spermaria) of the male; the former provide the ova, the latter the sperm. In the craniota we always find only one pair of gonads; in the amphioxus several pairs, arranged in succession. They must have had the same form in our hypothetical prospondylus (Figures 1.98 and 1.100 s). These segmental pairs of gonads are the original ventral halves of the coelom-pouches.