Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
One of the most unfortunate errors that this led to was the idea of an
original antithesis of germ and yelk. The latter was regarded as a
foreign body, extrinsic to the real germ, whereas it is properly a
part of it, an embryonic organ of nutrition. Many authors said there
was no trace of the embryo until a later stage, and outside the yelk;
sometimes the two-layered embryonic disk itself, at other times only
the central portion of it (as distinguished from the germinative area,
which we will describe presently), was taken to be the first outline
of the embryo. In the light of the gastraea theory it is hardly
necessary to dwell on the defects of this earlier view and the
erroneous conclusions drawn from it. In reality, the first
segmentation-cell, and even the stem-cell itself and all that issues
therefrom, belong to the embryo. As the large original yelk-mass in
the undivided egg of the bird only represents an inclosure in the
greatly enlarged ovum, so the later contents of its embryonic yelk-sac
(whether yet segmented or not) are only a part of the entoderm which
forms the primitive gut. This is clearly shown by the ova of the
amphibia and cyclostoma, which explain the transition from the
yelk-less ova of the amphioxus to the large yelk-filled ova of the
reptiles and birds.
It is precisely in the study of these difficult features that we see
the incalculable value of phylogenetic considerations in explaining
complex ontogenetic facts, and the need of separating cenogenetic
phenomena from palingenetic. This is particularly clear as regards the
comparative embryology of the vertebrates, because here the
phylogenetic unity of the stem has been already established by the
well-known facts of paleontology and comparative anatomy. If this
unity of the stem, on the basis of the amphioxus, were always borne in
mind, we should not have these errors constantly recurring.
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