Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
In order to get a clear idea of this, we must understand carefully the
relation of the embryonic shield to the germinative area and the
embryonic vesicle. This is done best by a comparison of the five
stages which are shown in longitudinal section in Figures 1.138 to
1.142. The embryonic shield (c), which at first projects very slightly
over the surface of the germinative area, soon begins to rise higher
above it, and to separate from the embryonic vesicle. At this point
the embryonic shield, looked at from the dorsal surface, shows still
the original simple sandal-shape (Figures 1.133 to 1.135). We do not
yet see any trace of articulation into head, neck, trunk, etc., or
limbs. But the embryonic shield has increased greatly in thickness,
especially in the anterior part. It now has the appearance of a thick,
oval swelling, strongly curved over the surface of the germinative
area. It begins to sever completely from the embryonic vesicle, with
which it is connected at the ventral surface. As this severance
proceeds, the back bends more and more; in proportion as the embryo
grows the embryonic vesicle decreases, and at last it merely hangs as
a small vesicle from the belly of the embryo (Figure 1.142 ds). In
consequence of the growth-movements which cause this severance, a
groove-shaped depression is formed at the surface of the vesicle, the
limiting furrow, which surrounds the vesicle in the shape of a pit,
and a circular mound or dam (Figure 1.139 ks) is formed at the outside
of this pit by the elevation of the contiguous parts of the germinal
vesicle.
(FIGURE 1.136. Transverse section of the embryonic disk of a chick at
the end of the first day of incubation, magnified about twenty times.
The edges of the medullary plate (m), the medullary swellings (w),
which separate the medullary from the horn-plate (h), are bending
towards each other. At each side of the chorda (ch) the primitive
segment plates (u) have separated from the lateral plates (sp). A
gut-gland layer. (From Remak.))
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