Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
(FIGURE 1.14. The human ovum, taken from the female ovary, magnified
500 times. The whole ovum is a simple round cell. The chief part of
the globular mass is formed by the nuclear yelk (deutoplasm), which is
evenly distributed in the active protoplasm, and consists of numbers
of fine yelk-granules. In the upper part of the yelk is the
transparent round germinal vesicle, which corresponds to the nucleus.
This encloses a darker granule, the germinal spot, which shows a
nucleolus. The globular yelk is surrounded by the thick transparent
germinal membrane (ovolemma, or zona pellucida). This is traversed by
numbers of lines as fine as hairs, which are directed radially towards
the centre of the ovum. These are called the pore-canals; it is
through these that the moving spermatozoa penetrate into the yelk at
impregnation.
FIGURE 1.15. A fertilised ovum from the oviduct of a hen. the yellow
yelk (c) consists of several concentric layers (d), and is enclosed in
a thin yelk-membrane (a). The nucleus or germinal vesicle is seen
above in the cicatrix or "tread" (b). From that point the white yelk
penetrates to the central yelk-cavity (d apostrophe). The two kinds of
yelk do not differ very much.
FIGURE 1.16. A creeping amoeba (highly magnified). The whole organism
is a simple naked cell, and moves about by means of the changing arms
which it thrusts out of and withdraws into its protoplasmic body.
Inside it is the roundish nucleus with its nucleolus.)
When the mature bird-ovum has left the ovary and been fertilised in
the oviduct, it covers itself with various membranes which are
secreted from the wall of the oviduct. First, the large clear
albuminous layer is deposited around the yellow yelk; afterwards, the
hard external shell, with a fine inner skin. All these gradually
forming envelopes and processes are of no importance in the formation
of the embryo; they serve merely for the protection of the original
simple ovum. We sometimes find extraordinarily large eggs with strong
envelopes in the case of other animals, such as fishes of the shark
type. Here, also, the ovum is originally of the same character as it
is in the mammal; it is a perfectly simple and naked cell. But, as in
the case of the bird, a considerable quantity of nutritive yelk is
accumulated inside the original yelk as food for the developing
embryo; and various coverings are formed round the egg. The ovum of
many other animals has the same internal and external features. They
have, however, only a physiological, not a morphological, importance;
they have no direct influence on the formation of the foetus. They are
partly consumed as food by the embryo, and partly serve as protective
envelopes. Hence we may leave them out of consideration altogether
here, and restrict ourselves to material points--TO THE SUBSTANTIAL
IDENTITY OF THE ORIGINAL OVUM IN MAN AND THE REST OF THE ANIMALS
(Figure 1.13).
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