Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The cutaneous glands are originally merely solid cone-shaped growths
of the epidermis, which sink into the underlying corium (Figure 2.286
1). Afterwards a canal (2, 3) is formed inside them, either by the
softening and dissolution of the central cells or by the secretion of
fluid internally. Some of the glands, such as the sudoriferous, do not
ramify (Figure 2.284 efg). These glands, which secrete the
perspiration, are very long, and have a spiral coil at the end, but
they never ramify; so also the wax-glands of the ears. Most of the
other cutaneous glands give out buds and ramify; thus, for instance,
the lachrymal glands of the upper eye-lid that secrete tears (Figure
2.286), and the sebaceous glands which secrete the fat in the skin and
generally open into the hair-follicles. Sudoriferous and sebaceous
glands are found only in mammals. But we find lachrymal glands in all
the three classes of Amniotes--reptiles, birds, and mammals. They are
wanting in the lower aquatic vertebrates.
(FIGURE 2.286. Rudimentary lachrymal glands from a human embryo of
four months. (From Kolliker.) 1 earliest structure, in the shape of a
simple solid cone, 2 and 3 more advanced structures, ramifying and
hollowing out. a solid buds, e cellular coat of the hollow buds, f
structure of the fibrous envelope, which afterwards forms the corium
about the glands.)
The mammary glands (Figures 2.287 and 2.288) are very remarkable; they
are found in all mammals, and in these alone. They secrete the milk
for the feeding of the new-born mammal. In spite of their unusual size
these structures are nothing more than large sebaceous glands in the
skin. The milk is formed by the liquefaction of the fatty milk-cells
inside the branching mammary-gland tubes (Figure 2.287 c), in the same
way as the skin-grease or hair-fat, by the solution of fatty cells
inside the sebaceous glands. The outlets of the mammary glands enlarge
and form sac-like mammary ducts (b); these narrow again (a), and open
in the teats or nipples of the breast by sixteen to twenty-four fine
apertures. The first structure of this large and elaborate gland is a
very simple cone in the epidermis, which penetrates into the corium
and ramifies. In the new-born infant it consists of twelve to eighteen
radiating lobes (Figure 2.288). These gradually ramify, their ducts
become hollow and larger, and rich masses of fat accumulate between
the lobes. Thus is formed the prominent female breast (mamma), on the
top of which rises the teat or nipple (mammilla). The latter is only
developed later on, when the mammary gland is fully-formed; and this
ontogenetic phenomenon is extremely interesting, because the earlier
mammals (the stem-forms of the whole class) have no teats. In them the
milk comes out through a flat portion of the ventral skin that is
pierced like a sieve, as we still find in the lowest living mammals,
the oviparous Monotremes of Australia. The young animal licks the milk
from the mother instead of sucking it.
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