Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
membrane," in the lower Vertebrates (Chapter 1.5). Underneath the
upper eye-lid are the lachrymal glands, the product of which, the
lachrymal fluid, keeps the outer surface of the eye smooth and clean.
Immediately under the sclerotic we find a very delicate, dark-red
membrane, very rich in blood-vessels--the choroid coat--and inside
this the retina (o), the expansion of the optic nerve (i). The latter
is the second cerebral nerve. It proceeds from the optic thalami (the
second cerebral vesicle) to the eye; penetrates its outer envelopes,
and then spreads out like a net between the choroid and the corpus
vitreum. Between the retina and the choroid there is a very delicate
membrane, which is usually (but wrongly) associated with the latter.
This is the black pigment-membrane (n). It consists of a single
stratum of graceful, hexagonal, regularly-joined cells, full of
granules of black colouring matter. This pigment membrane clothes, not
only the inner surface of the choroid proper, but also the hind
surface of its anterior muscular continuation, which covers the edge
of the lens in front as a circular membrane, and arrests the rays of
light at the sides. This is the well-known iris of the eye (h),
coloured differently in different individuals (blue, grey, brown,
etc.); it forms the anterior border of the choroid. The circular
opening that is left in the middle is the pupil, through which the
rays of light penetrate into the eye. At the point where the iris
leaves the anterior border of the choroid proper the latter is very
thick, and forms a delicate crown of folds (g), which surrounds the
edge of the lens with about seventy large and many smaller rays
(corona ciliaris.)
At a very early stage a couple of pear-shaped vesicles develop from
the foremost part of the first cerebral vesicle in the embryo of man
and the other Craniotes (Figures 1.155 a and 2.297 au). These growths
are the primary optic vesicles. They are at first directed outwards
and forwards, but presently grow downward, so that, after the complete
separation of the five cerebral vesicles, they lie at the base of the
intermediate brain. The inner cavities of these pear-shaped vesicles,
which soon attain a considerable size, are openly connected with the
ventricle of the intermediate brain by their hollow stems. They are
covered externally by the epidermis.
(FIGURE 2.316. Face of a human embryo, eight weeks old (From Ecker.))
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