(FIGURE 2.342. Skeleton of the hand or fore foot of six mammals. I man, II dog, III pig, IV ox, V tapir, VI horse. r radius, u ulna, a scaphoideum, b lunare, a triquetrum, d trapezium, e trapezoid, f capitatum, g hamatum, p pisiforme. 1 thumb, 2 index finger, 3 middle finger, 4 ring finger, 5 little finger. (From Gegenbaur.)) We have not space here to go into the details of Gegenbaur's theory of the skull. I must be content to refer the reader to the great work I have mentioned, in which it is thoroughly established from the empirico-philosophical point of view. He has also given a comprehensive and up-to-date treatment of the subject in his Comparative Anatomy of the Vertebrates (1898). Gegenbaur indicates as original "cranial ribs," or "lower arches of the cranial vertebrae," at each side of the head of the Selachii (Figure 2.334), the following pairs of arches: I and II, two lip-cartilages, the anterior (a) of which is composed of an upper piece only, the posterior (bc) from an upper and lower piece; III, the maxillary arches, also consisting of two pieces on each side--the primitive upper jaw (os palato-quadratum, o) and the primitive lower jaw (u); IV, the hyaloid bone (II); finally, V to X, six branchial arches in the narrower sense (III to VIII). From the anatomic features of these nine to ten cranial ribs or "lower vertebral arches" and the cranial nerves that spread over them, it is clear that the apparently simple cartilaginous primitive skull of the Selachii was originally formed from so many (at least nine) somites or provertebrae. The blending of these primitive segments into a single capsule is, however, so ancient that, in virtue of the law of curtailed heredity, the original division seems to have disappeared; in the embryonic development it is very difficult to detect it in isolated traces, and in some respects quite impossible. It is claimed that several (three to six) traces of provertebrae have been discovered in the anterior (pre-chordal) part of the Selachii-skull; this would bring up the number of cranial somites to twelve or sixteen, or even more. (FIGURES 2.343 TO 2.345. Arm and hand of three anthropoids. FIGURE 2.343. Chimpanzee (Anthropithecus niger). FIGURE 2.344. Veddah of Ceylon (Homo veddalis). FIGURE 2.345. European (Homo mediterraneus). (From Paul and Fritz Sarasin.))