Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
Underneath the stomach there now develops the disproportionately long
stretch of the small intestine. The development of this section is
very simple, and consists essentially in an extremely rapid and
considerable growth lengthways. It is at first very short, quite
straight, and simple. But immediately behind the stomach we find at an
early stage a horseshoe-shaped bend and loop of the gut, in connection
with the severance of the alimentary canal from the yelk-sac and the
development of the first mesentery. The thin delicate membrane that
fastens this loop to the ventral side of the vertebral column, and
fills the inner bend of the horseshoe formation, is the first rudiment
of the mesentery (Figure 1.147 g). We find at an early stage a
considerable growth of the small intestine; it is thus forced to coil
itself in a number of loops. The various sections that we have to
distinguish in it are differentiated in a very simple way--the
duodenum (next to the stomach), the succeeding long jejunum, and the
last section of the small intestine, the ileum.
From the duodenum are developed the two large glands that we have
already mentioned--the liver and pancreas. The liver appears first in
the shape of two small sacs, that are found to the right and left
immediately behind the stomach (Figures 2.353 f, and 2.354 c). In many
of the lower Vertebrates they remain separate for a long time (in the
Myxinoides throughout life), or are only imperfectly joined. In the
higher Vertebrates they soon blend more or less completely to form a
single large organ. The growth of the liver is very brisk at first. In
the human embryo it grows so much in the second month of development
that in the third it occupies by far the greater part of the
body-cavity (Figure 2.357). At first the two halves develop equally;
afterwards the left falls far behind the right. In consequence of the
unsymmetrical development and turning of the stomach and other
abdominal viscera, the whole liver is now pushed to the right side.
Although the liver does not afterwards grow so disproportionately, it
is comparatively larger in the embryo at the end of pregnancy than in
the adult. Its weight relatively to that of the whole body is 1 : 36
in the adult, and 1 : 18 in the embryo. Hence it is very important
physiologically during embryonic life; it is chiefly concerned in the
formation of blood, not so much in the secretion of bile.
Immediately behind the liver a second large visceral gland develops
from the duodenum, the pancreas or sweetbread. It is wanting in most
of the lowest classes of Vertebrates, and is first found in the
fishes. This organ is also an outgrowth from the gut.
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