Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
Our phylogenetic interpretation of the ovum, and the reduction of it
to some ancient amoeboid ancestral form, supply the answer to the old
problem: "Which was first, the egg or the chick?" We can now give a
very plain answer to this riddle, with which our opponents have often
tried to drive us into a corner. The egg came a long time before the
chick. We do not mean, of course, that the egg existed from the first
as a bird's egg, but as an indifferent amoeboid cell of the simplest
character. The egg lived for thousands of years as an independent
unicellular organism, the Amoeba. The egg, in the modern physiological
sense of the word, did not make its appearance until the descendants
of the unicellular Protozoon had developed into multicellular animals,
and these had undergone sexual differentiation. Even then the egg was
first a gastraea-egg, then a platode-egg, then a vermalia-egg, and
chordonia-egg; later still acrania-egg, then fish-egg, amphibia-egg,
reptile-egg, and finally bird's egg. The bird's egg we have experience
of daily is a highly complicated historical product, the result of
countless hereditary processes that have taken place in the course of
millions of years.
The earliest ancestors of our race were simple Protophyta, and from
these our protozoic ancestors were developed afterwards. From the
morphological point of view both the vegetal and the animal Protists
were simple organisms, individualities of the first order, or
plastids. All our later ancestors are complex organisms, or
individualities of a higher order--social aggregations of a plurality
of cells. The earliest of these, the Moraeada, which represent the
third stage in our genealogy, are very simple associations of
homogeneous, indifferent cells--undifferentiated colonies of social
Amoebae or Infusoria. To understand the nature and origin of these
protozoa-colonies we need only follow step by step the first embryonic
products of the stem-cell. In all the Metazoa the first embryonic
process is the repeated cleavage of the stem-cell, or first
segmentation-cell (Figure 2.229). We have already fully considered
this process, and found that all the different forms of it may be
reduced to one type, the original equal or primordial segmentation
(cf. Chapter 1.8). In the genealogical tree of the Vertebrates this
palingenetic form of segmentation has been preserved in the Amphioxus
alone, all the other Vertebrates having cenogenetically modified forms
of cleavage. In any case, the latter were developed from the former,
and so the segmentation of the ovum in the Amphioxus has a great
interest for us (cf. Figure 1.38). The outcome of this repeated
cleavage is the formation of a round cluster of cells, composed of
homogeneous, indifferent cells of the simplest character (Figure
2.230). This is called the morula (= mulberry-embryo) on account of
its resemblance to a mulberry or blackberry.
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