Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
Some of the infusoria of the flagellata-class (Signura, Magosphaera,
etc.) are similar in structure to these vegetal clusters, but differ
in their animal nutrition; they form the special group of the
Catallacta. In September, 1869, I studied the development of one of
these graceful animals on the island of Gis-Oe, off the coast of
Norway (Magosphaera planula), Figures 2.231 and 2.232). The
fully-formed body is a gelatinous ball, with its wall composed of
thirty-two to sixty-four ciliated cells; it swims about freely in the
sea. After reaching maturity the community is dissolved. Each cell
then lives independently for some time, grows, and changes into a
creeping amoeba. This afterwards contracts, and clothes itself with a
structureless membrane. The cell then looks just like an ordinary
animal ovum. When it has been in this condition for some time the cell
divides into two, four, eight, sixteen, thirty-two, and sixty-four
cells. These arrange themselves in a round vesicle, thrust out
vibratory lashes, burst the capsule, and swim about in the same
magosphaera-form with which we started. This completes the life-circle
of the remarkable and instructive animal.
If we compare these permanent blastulae with the free-swimming
ciliated larvae or blastulae, with similar construction, of many of
the lower animals, we can confidently deduce from them that there was
a very early and long-extinct common stem-form of substantially the
same structure as the blastula. We may call it the Blastaea. Its body
consisted, when fully formed, of a simple hollow ball, filled with
fluid or structureless jelly, with a wall composed of a single stratum
of ciliated cells. There were probably many genera and species of
these blastaeads in the Laurentian period, forming a special class of
marine protists.
It is an interesting fact that in the plant kingdom also the simple
hollow sphere is found to be an elementary form of the multicellular
organism. At the surface and below the surface (down to a depth of
2000 yards) of the sea there are green globules swimming about, with a
wall composed of a single layer of chlorophyll-bearing cells. The
botanist Schmitz gave them the name of Halosphaera viridis in 1879.
The next stage to the Blastaea, and the sixth in our genealogical
tree, is the Gastraea that is developed from it. As we have already
seen, this ancestral form is particularly important. That it once
existed is proved with certainty by the gastrula, which we find
temporarily in the ontogenesis of all the Metazoa (Figure 1.29 J, K).
As we saw, the original, palingenetic form of the gastrula is a round
or oval uni-axial body, the simple cavity of which (the primitive gut)
has an aperture at one pole of its axis (the primitive mouth). The
wall of the gut consists of two strata of cells, and these are the
primary germinal layers, the animal skin-layer (ectoderm) and vegetal
gut-layer (entoderm).
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