Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
on this account it has recently been regarded as the representative of
a special order of gastraeads (Mesogastria).
Probably a near relative of the Pemmatodiscus is the Kunstleria
Gruveli (Figure 2.233, 2). It lives in the body-cavity of Vermalia
(Sipunculida), and differs from the former in having no lashes either
on the large ectodermic cells (e) or the small entodermic (i); the
germinal layers are separated by a thick, cup-shaped, gelatinous mass,
which has been called the "clear vesicle" (f). The primitive mouth is
surrounded by a dark ring that bears very strong and long vibratory
lashes, and effects the swimming movements.
Pemmatodiscus and Kunstleria may be included in the family of the
Gastremaria. To these gastraeads with open gut are closely related the
Orthonectida (Rhopalura, Figure 2.233, 3 to 5). They live
parasitically in the body-cavity of echinoderms (Ophiura) and
vermalia; they are distinguished by the fact that their primitive
gut-cavity is not empty, but filled with entodermic cells, from which
the sexual cells are developed. These gastraeads are of both sexes,
the male (Figure 1.3) being smaller and of a somewhat different shape
from the oval female (Figure 1.4).
The somewhat similar Dicyemida (Figure 1.6) are distinguished from the
preceding by the fact that their primitive gut-cavity is occupied by a
single large entodermic cell instead of a crowded group of sexual
cells. This cell does not yield sexual products, but afterwards
divides into a number of cells (spores), each of which, without being
impregnated, grows into a small embryo. The Dicyemida live
parasitically in the body-cavity, especially the renal cavities, of
the cuttle-fishes. They fall in several genera, some of which are
characterised by the possession of special polar cells; the body is
sometimes roundish, oval, or club-shaped, at other times long and
cylindrical. The genus Conocyema (Figures 1.7 to 1.15) differs from
the ordinary Dicyema in having four polar pimples in the form of a
cross, which may be incipient tentacles.
The classification of the Cyemaria is much disputed; sometimes they
are held to be parasitic infusoria (like the Opalina), sometimes
platodes or vermalia, related to the suctorial worms or rotifers, but
having degenerated through parasitism. I adhere to the
phylogenetically important theory that I advanced in 1876, that we
have here real gastraeads, primitive survivors of the common
stem-group of all the Metazoa. In the struggle for life they have
found shelter in the body-cavity of other animals.
Public-domain text, read in full here on John Shaqi.
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