Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The Gastraea bilateralis, of which we may conceive the bilateral
gastrula of the amphioxus to be a palingenetic reproduction,
represented the two-sided organism of the earliest Metazoa in its
simplest form. The vegetal entoderm that lined their simple gut-cavity
served for nutrition; the ciliated ectoderm that formed the external
skin attended to locomotion and sensation; finally, the two primitive
mesodermic cells, that lay to the right and left at the ventral border
of the primitive mouth, were sexual cells, and effected reproduction.
In order to understand the further development of the gastraea, we
must pay particular attention to: (1) the careful study of the
embryonic stages of the amphioxus that lie between the gastrula and
the chordula; (2) the morphological study of the simplest Platodes
(Platodaria and Turbellaria) and several groups of unarticulated
Vermalia (Gastrotricha, Nemertina, Enteropneusta).
We have to consider the Platodes first, because they are on the border
between the two principal groups of the Metazoa, the Coelenteria and
the Coelomaria. With the former they share the lack of body-cavity,
anus, and vascular system; with the latter they have in common the
bilateral type, the possession of a pair of nephridia or renal canals,
and the formation of a vertical brain or cerebral ganglion. It is now
usual to distinguish four classes of Platodes: the two free-living
classes of the primitive worms (Platodaria) and the coiled-worms
(Turbellaria), and the two parasitic classes of the suctorial worms
(Trematoda) and the tape-worms (Cestoda). We have only to consider the
first two of these classes; the other two are parasites, and have
descended from the former by adaptation to parasitic habits and
consequent degeneration.
(FIGURE 2.239. Aphanostomum Langii (Haeckel), a primitive worm of the
platodaria class, of the order of Cryptocoela or Acoela. This new
species of the genus Aphanostomum, named after Professor Arnold Lang
of Zurich, was found in September, 1899, at Ajaccio in Corsica
(creeping between fucoidea). It is one-twelfth of an inch long,
one-twenty-fifth of an inch broad, and violet in colour. a mouth, g
auditory vesicle, e ectoderm, i entoderm, o ovaries, a spermaries, f
female aperture, m male aperture.)
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