Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
Next to the ancient stem-group of the Turbellaria come a number of
more recent chordonia ancestors, which we class with the Vermalia or
Helminthes, the unarticulated worms. These true worms (Vermes, lately
also called Scolecida) are the difficulty or the lumber-room of the
zoological classifier, because the various classes have very
complicated relations to the lower Platodes on the one hand and the
more advanced animals on the other. But if we exclude the Platodes and
the Annelids from this stem, we find a fairly satisfactory unity of
organisation in the remaining classes. Among these worms we find some
important forms that show considerable advance in organisation from
the platode to the chordonia stage. Three of these phenomena are
particularly instructive: (1) The formation of a true (secondary)
body-cavity (coeloma); (2) the formation of a second aperture of the
gut, the anus; and (3) the formation of a vascular system. The great
majority of the Vermalia have these three features, and they are all
wanting in the Platodes; in the rest of the worms at least one or two
of them are developed.
Next and very close to the Platodes we have the Ichthydina
(Gastrotricha), little marine and fresh-water worms, about 1/250 to
1/1000 inch long. Zoologists differ as to their position in
classification. In my opinion, they approach very close to the
Rhabdocoela (Figures 2.240 and 2.241), and differ from them chiefly in
the possession of an anus at the posterior end (Figure 2.242 a).
Further, the cilia that cover the whole surface of the Turbellaria are
confined in the Gastrotricha to two ciliated bands (f) on the ventral
surface of the oval body, the dorsal surface having bristles.
Otherwise the organisation of the two classes is the same. In both the
gut consists of a muscular gullet (s) and a glandular primitive gut
(d). Over the gullet is a double brain (acroganglion, g). At the side
of the gut are two serpentine prorenal canals (water-vessels or
pronephridia, nc), which open on the ventral side (nm). Behind are a
pair of simple sexual glands or gonads (Figure 2.243 e).
While the Ichthydina are thus closely related to the Platodes, we have
to go farther away for the two classes of Vermalia which we unite in
the group of the "snout-worms" (Frontonia). These are the Nemertina
and the Enteropneusta. Both classes have a complete ciliary coat on
the epidermis, a heritage from the Turbellaria and the Gastraeads;
also, both have two openings of the gut, the mouth and anus, like the
Gastrotricha. But we find also an important organ that is wanting in
the preceding forms--the vascular system. In their more advanced
mesoderm we find a few contractile longitudinal canals which force the
blood through the body by their contractions; these are the first
blood-vessels.
(FIGURE 2.244. A simple Nemertine. m mouth, d gut, a anus, g brain, n
nerves, h ciliary coat, ss sensory pits (head-clefts), au eyes, r
dorsal vessel, l lateral vessels. (Diagram.)
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