Anatomy, Comparative; Embryology, Human; Evolution; Human beings -- Origin
The instructive group of the Permian Tocosauria, the common root from
which the divergent stems of the Sauropsids and mammals have issued,
merits our particular attention as the stem-group of all the Amniotes.
Fortunately a living representative of this extinct ancestral group
has been preserved to our day; this is the remarkable lizard of New
Zealand, Hatteria punctata (Figure 2.264). Externally it differs
little from the ordinary lizard; but in many important points of
internal structure, especially in the primitive construction of the
vertebral column, the skull, and the limbs, it occupies a much lower
position, and approaches its amphibian ancestors, the Stegocephala.
Hence Hatteria is the phylogenetically oldest of all living reptiles,
an isolated survivor from the Permian period, closely resembling the
common ancestor of the Amniotes. It must differ so little from this
extinct form, our hypothetical Protamniote, that we put it next to the
Proreptilia. The remarkable Permian Palaehatteria, that Credner
discovered in the Plauen terrain at Dresden in 1888, belongs to the
same group (Figure 2.266). The Jurassic genus Homoeosaurus (Figure
2.265), of which well-preserved skeletons are found in the Solenhofen
schists, is perhaps still more closely related to them.
Unfortunately, the numerous fossil remains of Permian and Triassic
Tocosauria that we have found in the last two decades are, for the
most part, very imperfectly preserved. Very often we can make only
precarious inferences from these skeletal fragments as to the anatomic
characters of the soft parts that went with the bony skeleton of the
extinct Tocosauria. Hence it has not yet been possible to arrange
these important fossils with any confidence in the ancestral series
that descend from the Protamniotes to the Sauropsids on the one side
and the Mammals on the other. Opinions are particularly divided as to
the place in classification and the phylogenetic significance of the
remarkable Theromorpha. Cope gives this name to a very interesting and
extensive group of extinct terrestrial reptiles, of which we have only
fossil remains from the Permian and Triassic strata. Forty years ago
some of these Therosauria (fresh-water animals) were described by Owen
as Anomodontia. But during the last twenty years the distinguished
American paleontologists, Cope and Osborn, have greatly increased our
knowledge of them, and have claimed that the stem-forms of the Mammals
must be sought in this order. As a matter of fact, the Theromorpha are
nearer to the Mammals in the chief points of structure than any other
reptiles. This is especially true of the Thereodontia, to which the
Pureosauria and Pelycosauria belong (Figure 2.267). The whole
structure of their pelvis and hind-feet has attained the same form as
in the Monotremes, the lowest Mammals. The formation of the scapula
and the quadrate bone shows an approach to the Mammals such as we find
in no other group of reptiles.
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