The marshes of European countries, and also those of warmer lands,
often contain bladderworts, or Utriculariæ (Fig. 23)--floating
water-plants, without roots, and with horizontally spread,
long-drawn-out, tendril-like shoots, in part thickly covered with
whorls of delicate, needle-shaped leaves, in part bearing sparse
leaves of quite peculiar structure. These are stalked, hollow
bladders (Fig. 23 _A_, _FA_), with quite a narrow entrance at the
apex, which is closed, as far as larger animals are concerned, by
projecting bristle-like hairs (_B_). Small animals, such as water-fleas
(_Daphnia_), species of _Cyclops_, and Ostracods, can swim in between
the bristles, and they then come in contact with a valve which opens
easily inwards (_B_, _kl_) and allows them to penetrate into the
interior of the trap. Once inside they are captives, for the valve
does not open outwards; therefore they soon die and decompose, and are
then taken up by special absorptive cells (_B_, _C_, _sz_) and utilized
as nourishment for the plants. In this way the Utriculariæ catch
numerous little crustaceans and insect larvæ, which slip into their
traps, presumably for concealment.
[Illustration: FIG. 24. Pitcher of _Nepenthes villosa_, after Kerner.
_St_, stalk of the leaf. _Spr_, its apex. _Fk_, the pitcher. _R_, the
margin beset with incurved spines.]
Another example is found in the marsh plants of the genus _Nepenthes_,
some species of which live as climbers on the outskirts of tropical
forests, climbing up the trees and letting their long, thin tendrils
hang downwards, often over ponds and stagnant pools, where swarms of
small flying insects abound. These plants have developed exceedingly
remarkable contrivances for catching insects and using them as food
(Fig. 24). The long stalks (_St_) of their leaves (_Spr_) are first
bent downwards, then they suddenly turn sharply upwards, and the
upturned portion is modified into a pitcher-like structure, in the
bottom of which a fluid gathers, acid in taste, containing pepsin,
and therefore a digestive fluid. Nitrogenous substances, such as
flesh, dissolve in this fluid, and insects which fall into the pitcher
from the rim are killed and dissolved. There are many species of
_Nepenthes_, but not all of them possess the trap-structure in equal
perfection, so that we are able, to some extent, to follow the course
of its evolution, from a broad leaf-stalk, somewhat bent over at the
edges, to the marvellous closed pitcher shown by _Nepenthes villosa_
(Fig. 24) of Borneo. In this species the pitchers attain a length of
fifty centimetres, and are beautifully coloured, resembling in that
respect, as well as in their form, the tobacco-pipe-like flowers of
the tropical Aristolochiæ. When we come to discuss the origin of
flowers, we shall see that the bright, conspicuous colour possesses
a very considerable value in attracting insects; and in the case of
the pitcher-plant, too, the gorgeous colour probably allures insects
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