This is best shown by the fact that the instincts may lead their
possessors astray, which always happens when an animal is transferred
to an unnatural situation, to which its instincts are not adapted, so
to speak. The mole-cricket, which is in the habit of escaping pursuit
by burrowing in the earth, makes violent motions with the forelegs,
even if it be placed upon a plate of glass into which it could not
possibly burrow; an ant-lion (_Myrmeleo_), whose instinct impels
it to bore into loose sand by pushing backwards with the abdomen,
goes backwards on a plate of glass as soon as danger threatens, and
endeavours, with the utmost exertions, to bore into it. It knows no
other mode of flight, and its intellect is much too weak to suggest any
novel mode. Even the mode of escape most universal among animals, that
of simply running away, does not occur to it; it acts as it must in
accordance with its inborn instinct; it cannot do otherwise.
The change of instincts in the different stages of development of
one and the same animal have always seemed to me very remarkable; for
instance, the change of the food-instinct in the caterpillar and the
butterfly, where the food-instinct is liberated in the caterpillar
by the leaf of a particular plant, but in the butterfly by the sight
and fragrance of a flower, the nectar of which it sucks. In this case
everything is different in the two stages of development, the whole
apparatus for seeking and taking food, as well as the nerve-mechanism
which determines these modes of action. And how far apart often are the
stimuli which liberate the instinct! The larva of the flower-visiting,
honey-sucking _Eristalis tenax_ is the ugly, white, so-called
rat-tailed larva, well described by Réaumur, which lives swimming
in liquid manure, and feeds on that! What complete and far-reaching
changes, not only in the visible structure, but also in the finer
nervous mechanisms, which we cannot yet verify, must have taken place
in the vicissitudes of time and circumstance during the life-history of
this insect!
[Illustration: FIG. 32. Metamorphosis of _Sitaris humeralis_, an
oil-beetle, after Fabre. _a_, first larval form, much enlarged. _b_,
second larval form. _c_, resting stage of this larva (so-called
'pseudo-pupa'). _d_, third larval form. _e_, pupa.]
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