The biting mouth-parts of beetles, Neuroptera, and Orthoptera (Fig.
48), consist of three pairs of jaws, of which the first, the mandibles
(_md_), are simply powerful pincers for seizing and tearing or chewing
the food. They have no part in the development of the suctorial
apparatus either in bees or in butterflies, so they may be left out
of account. The two other pairs of jaws, the first and second maxillæ
(_mx_^1 and _mx_^2), are constructed exactly on the same type, having
a jointed basal portion (_st_) bearing two lobes, an external (_le_)
and an internal (_li_), and a feeler or palp, usually with several
joints, directed outwards from the lobes (_pm_ and _pl_). The second
pair of maxillæ (_mx_^2) differs from the first chiefly in this, that
the components of the pair meet in the median line of the body, and
fuse more or less to form the so-called 'under lip' or labium. In the
example given, the cockroach (_Periplaneta orientalis_), this fusion
is only partial, the lobes having remained separate (_le_ and _li_);
and the same is true of the bee, but in this case the inner lobes have
grown into a long worm-like process which is thrust into the nectar in
the act of sucking.
[Illustration: FIG. 49. Head of the Bee. _Au_, compound eyes. _au_,
ocelli. _at_, antennæ. _la_, upper lip. _md_, mandibles. _mx_^1, first
maxillæ, with _pm_, the rudimentary maxillary palp. _mx_^2, second
maxillæ with the internal lobes (_li_) fused to form the 'tongue.'
_le_, the external lobes of the second maxillæ, known as 'paraglossæ.'
_pl_, labial palp.]
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