Thus the corolla of many flowers is suited to the thick, hairy head
and thorax of the bee, for to these only does the pollen adhere in
sufficient quantity to fertilize the next flower; yet the same flowers
are frequently visited by butterflies, and in many of them there
has been no adaptation to prevent these useless visits. Obviously
this is because preventive arrangements could only begin, according
to our theory, when they were necessary to the preservation of the
species; in this case, therefore, only when the pillaging visits
of the butterflies withdrew so many flowers from the influence of
the effective pollinating visitor, the bee, that too few seeds were
formed, and the survival of the species was threatened by the continual
dwindling of the normal number. As long as the bees visit the flowers
frequently enough to ensure the formation of the necessary number of
seeds a process of selection could not set in; but should the bees
find, for instance, that nearly all the flowers had been robbed of
their nectar, and should therefore visit them less diligently, then
every variation of the flower which made honey less accessible to the
butterflies would become the objective of a process of selection.
Everywhere we find similar imperfections of adaptation which indicate
that they must depend on processes of selection. Thus numerous flowers
are visited by insects other than those which pollinate them, and these
bring them no advantage, but merely rob them of nectar and pollen;
the most beautiful contrivances of many flowers, such as _Glycinia_,
which are directed towards cross-fertilization by bees, are rendered
of no effect because wood-bees and humble-bees bite holes into the
nectaries from the outside, and so reach the nectar by the shortest
way. I do not know whether bees in the native land of the _Glycinia_
do the same thing, but in any case they can do no sensible injury to
the species, since otherwise processes of selection would have set in
which would have prevented the damage in some way or other, whether by
the production of stinging-hairs, or hairs with a burning secretion, or
in some other way. If the actual constitution of the plant made this
impossible, the species would become less abundant and would gradually
die out.
Thus the relative imperfection of the flower-adaptations, which in
general are so worthy of admiration, affords a further indication that
their origin is due to processes of selection.
ADDITIONAL NOTE TO CHAPTER X.
Public-domain text, read in full here on John Shaqi.
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