the lime-tree as they do on the deep-blue gentian of the Alpine meadows
or the red blossoms of the Weigelia, the dog-roses of our gardens or
the yellow buttercups (_Ranunculus_) of our meadows; they despise
nothing that yields them honey. But it certainly does not follow
from this that the bees may not, under certain circumstances, have
exercised a selecting influence upon the fixation and intensification
of a new colour-variety of a flower. This is less a question of a
_colour-preference_, in the human sense, on the part of the bees
than of the _greater visibility_ of the colour in question in the
environment peculiar to the flower, and of the amount of rivalry the
bees meet with from other insects in regard to the same flower. In
individual cases this would be difficult to demonstrate, especially
since we can form only an approximate idea of the insect's power of
seeing colour, and cannot judge what the colours of the individual
blossoms count for in the mosaic picture of a flowery meadow. Yet this
is the important point, for, as soon as the bees perceive one colour
more readily than another, the preponderance of this colour-variety
over other variations is assured, since it will be more frequently
visited. In the same way we cannot guess in individual cases why one
species of flower should exhale perfume while a nearly related species
does not. But when we remember that many flowers adapted for the visits
of dipterous insects possess a nauseous carrion-like smell, by means
of which they not only attract flies but scare off other insects, we
can readily imagine cases in which it was of importance to a flower
to be able to be easily found by bees without betraying itself by its
pleasant fragrance to other less desirable visitors.
Thus, therefore, we can understand the odourless but intensely blue
species of gentian, if we may assume that its blue colour is more
visible to bees than to other insects. If I were to elaborate in
detail all the principles which here suggest themselves to me I should
require to write a complete section, and I am unwilling to do this
until I can bring forward a much larger number of new observations
than I am at present in a position to do. All I wish to do here is
to exhort doubters to modesty, and to remind them that these matters
are exceedingly complex, and that we should be glad and grateful that
expert observers like Darwin and Hermann Müller have given us some
insight into the principles interconnecting the facts, instead of
imagining whenever we meet with some little apparently contradictory
fact, which may indeed be quite correct in itself, that the whole
theory of the development of flowers through insects has been
overthrown. Let us rather endeavour to understand such facts, and to
arrange them in their places as stones of the new building.
Public-domain text, read in full here on John Shaqi.
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