It can be proved, however, that such transmissions may, in the
course of many successive generations, become intensified until the
characters are exhibited by the females in the same degree as in the
males. I know no better example of this than that afforded by the
beautiful butterflies of the genus _Lycæna_. In this genus, which is
rich in species and widely distributed over the whole earth, and must
therefore be an old one, the upper surface of the wing is blue in by
far the greater number of species, at least in the male sex. But there
are three or four species which are dark-brown, and quite or nearly
alike in the two sexes; such are the species _Lycæna agestis_, _L.
eumedon_, _L. admetus_, and others. Everything indicates that this is
the primitive colour of the genus. Moreover, there are some species
with brown females, in which the males are not completely blue, but
which have a slight bluish tinge, like _L. alsus_, the smallest of
our indigenous Blues. Then follows a host of beautiful species, like
_L. alexis_, _L. adonis_, _L. damon_, _L. corydon_, and many others,
with brown females, and among these there occasionally occur females
more or less tinged with blue. These lead on to _L. meleager_, which
has two forms of female, a common brown and a rarer blue; and thus
we reach _L. tiresias_, _L. optilete_, and _L. argiolus_, in which
all the females are blue, although less intensely and completely so
than their mates. The climax of this evolutionary series is reached
by some species like _L. beatica_, belonging to tropical or at least
warm countries, in which both sexes are of an equally intense blue. As
we know that, in species with an excess of males, sexual characters
always begin in the males, there can be no doubt as to the direction
of evolution--from brown to blue--in this series. Furthermore, the
entire absence of scent-scales in most of the species with brown males
indicates the great age of these species, for, as far as I have been
able to investigate, all the males of the blue species possess them.
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